Accepted Name - Basionymum / Basionym - Synonyms - Ind. loc. [Indicatio locotypica (type locality)] - Lectotypus [Lectotype] - Designated by - Host - Comments - Chromosomatic number - Ic. [Images] - Distribution - Specimens Examined - References - Authors - Cite us

ISSN: 2386-9666 - Last Updated: 17 June 2024


Accepted Name:

Cistanche flava (C.A. Mey.) Korsh. in Zap. Imp. Akad. Nauk Fiz.-Mat. Otd.: 5 (1896)

Basionymum / Basionym:

Phelypaea flava C.A. Mey. in Verz. Pfl. Casp. Meer.: 104 (1831 [Nov-Dec 1831])

Synonyms:

Homotypic synonyms:

Cistanche flava O. Fedtsch. & B. Fedtsch., Consp. Fl. Turkestanicae 5: 112 (1913), isonym? (https://ipni.org/n/661954-1 - https://www.gbif.org/es/species/7332238)

Cistanche tubulosa var. flava (C.A. Mey.) Beck in Bull. Herb. Boissier 2(4): 686 (1904) (Beck 1930: 33)

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Heterotypic synonyms:

? = Phelypaea gigantea Kar. in Bull. Soc. Imp. Naturalistes Mosc. 1839(2): 165 [no. 638] (1839), nom. nud. (Ledebour 1849: 314, sub P. flava C. A. Meyer Ind. cauc. p. 104, "Karel. Enum. pl. Turkomann. in Bullet. de la  Soc. des Natur. d. Mosc. p. a. 1839, p. 165, No. 638"; Boissier 1879: 501, sub P. flava; Beck 1930: 36, sub C. salsa; Schlauer & al. 2003; Beck 1930: 33, sub C. tubulosa var. flava; Novopokrovsky & Tzvelev 1958: 23[21], sub C. flava). Ind. loc.: Turkmenistan: "In insula Ogurtsch'nsk [Ogurtschinsky Island / Ogurchinsky / Ogurja Ada / Aidak Island in the sutheastern part of Caspian Sea]" (Karelin 1839: 165). "Siberia uralensi in insula Ogurtschik [Ogurtschinsky Island / Ogurchinsky / Ogurja Ada / Aidak Island] m. caspii [Caspian Sea]! (Karel)" (Ledebour 1849: 314, sub P. flava). "auf der Insel Ogurtschinsk am Caspi See (Karelin als Ph. gigantea!)" (Beck 1930: 37, sub C. salsa)

Phelypaea tubulosa Schenk, Pl. Spec. Schubert: 23[-24] (1840 [1 Jun-1 Dec 1840?]) ["138. Ph. tubulosa n. sp."] (Walpers 1844-1845: 462, sub P. tubulosa; Reuter 1847: 12, sub P. tubulosa; Hooker f. 1884: 324; Beck 1930: 32, sub C. tubulosa; Boulos 2002: 94; Moreno Moral, Sánchez Pedraja & Piwowarczyk 2018: 27). Syn.: Cistanche tubulosa (Schenk) Hook. f., Fl. Brit. Ind. 4(11): 324 (1884); = Cistanche tinctoria (Forssk.) Deflers in Bull. Soc. Bot. France 43: 327 no. 336 (1896) (Moreno Moral, Sánchez Pedraja & Piwowarczyk 2018: 24). Ind. loc.: Egypt: "Hab. ad viam inter Suez et Tor [Sinai] in Arabia petraea, ubi legerunt Dr. Roth et Erdl mense Februario" (Schenk 1840: 24); "Inter Suez et Tor [Sinai], in Arabiâ Petraeâ, in radicibus..." (Walpers 1844-1845: 462, sub P. tubulosa; Reuter 1847: 12, sub P. tubulosa); "The type ... was collected in Sinai, between Suez and el-Tor [Egypt] by Roth & Erdl" (Boulos 2002: 95). Obs. "... Bracteae ternae, duabus lateralibus fere dimidio brevioribus, media maxima, calycem superante; ... Corolla ... tubo longissimo calycen duplo superante, arcuato-infundibuliformi ... Antherae ... obtusae ... villosos-lanatae ..." (Schenk 1840: 24). Type: Egypt, Sinai Peninsula: inter Suez et Tor in Arabia petraea, leg. Roth & Erdl (LMO s.n.) (Novopokrovsky & Tzvelev 1958: 23[21], sub C. tubulosa; Moreno Moral, Sánchez Pedraja & Piwowarczyk 2018: 27, sub C. tubulosa); K ? (TL-2: 283).

Cistanche tubulosa (Schenk) Hook. f., Fl. Brit. Ind. 4(11): 324 (1884) (Beck 1930: 33; Schlauer & al. 2002). Syn.: Phelypaea tubulosa Schenk, Pl. Spec. Schubert: 23[-24] (1840 [1 Jun-1 Dec 1840?]); = Cistanche tinctoria (Forssk.) Deflers in Bull. Soc. Bot. France 43: 327 no. 336 (1896) (Moreno Moral, Sánchez Pedraja & Piwowarczyk 2018: 24).

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= Cistanche flava subsp. bicolor Saeidi & Shahi, Iran. J. Bot. 14(2): 96 f.1 (2008 [31 Dec 2008]). Herbarium Specimens: "Type: Iran, Semnan, Touran protected area, 3 km from Chah-jam to Toroud, 1100 m, 1978.4.23, Freitag & Mozaffarian 28437 (holo. TARI)" (Saeidi & Shahi 2008: 96). Obs. "corolla color — yellow with violet limb ... Anther apex — mucronate" (Saeidi & Shahi 2008: 96)

? = Cistanche flava subsp. brevibracteata Ataei in Molec. Syst. Evol. Cistanche: 87-88 (2017) [unpublished thesis is not effective publication (Turland & al. 2018: Art. 29-31 (Art. 6.1))]. Herbarium Specimens: "Type: Iran. Southern Khorassan: 40-42 km after Nehbandan towards Zabol, 31°29′29''N, 60°36′34''E, 807-825 m, 30 March 2011, Akhani, Noormohammadi & Samadi 22074 (holotype TUH [n.v.])"; 1. "Davis & Bokhari (D.55976) / Cistanche tubulosa (Schenk) Wight / Iran. D6 (Times Atlas): Mond river / N of Kaki. 100 m. Lowe sand / dunes on S. bank. On Haloxylon. / Flowers yellow (bud pale violet) / 27 March 1974". – 2. "Cistanche flava  (C.A. Mey.) Korsh. subsp. brevibracteata / subsp. nov. / Det.: Najibeh Ataei May 2014 / ..." (https://data.rbge.org.uk/herb/E00028866 - E 00028866.zip - https://www.gbif.org/occurrence/575300308 (Ataei 2017: 88); rev. Ó. Sánchez Pedraja Jun 2024, bracteoles oblong-lanceolate shorter than calyx with scarious margins, corolla lobes tinged purple, anthers obtuse, see Cistanche tubulosa (Schenk) Hook. f.)); "Cistanche tubulosa (Schenk) R. Wight / Touran Protected Area [Khar Turan National Park, Semnan prov., Iran] / SE of Shahrud at road / to 1,5 km SW of Chejam / sand fields, Haloxylon persicum ass. / Alt.: 1100 m. / 23.04.1978 / Coll. H. Freitag 14695" (https://herbarium.bgbm.org/object/B100264808 - https://image.bgbm.org/images/internal/HerbarThumbs/B100264808_1700 - https://www.gbif.org/occurrence/911748649 (Ataei 2017: 88); rev. Ó. Sánchez Pedraja Jun 2024, bracteoles oblong-lanceolate shorter than calyx, corolla lobes tinged purple, anthers obtuse, see Cistanche tubulosa (Schenk) Hook. f.). Obs. "usually without scarious margins (1.1.8 C. flava)" (Ataei 2017: 62)

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Misapplied names:

Cistanche tubulosa auct., non Cistanche tubulosa (Schenk) Hook. f., Fl. Brit. Ind. 4(11): 324 (1884) (Beck 1930: 33; Schlauer & al. 2002)

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Ind. loc.: Azerbaijan: "Hab. in provinciis caucasicis [in arena mobili versus m. casp. inter Baku et Sallian pr. Karawauserai [Karvansaray / Karabansarai] Eszek ! (C.A. Meyer)]" (Ledebour 1849: 314, sub P. flava). "Hab. in arenâ mobili ad littus Caspium ad Erzek inter Baku et Sallian (C.A. Mey.!)" (Boissier 1879: 501, sub P. flava). "An den Küsten des Caspischen Meeres im Dünensande bei Erzek zwischen Baku und Sallian (C.A. Meyer)" (Beck 1930: 33). "Described from Azerbaidzhan [Azerbaijan] (between Baku and Salyans [Saljanami / Sallian])" (Novopokrovsky & Tzvelev 1958: 24[22], sub C. flava).

Typus / Type: «Lectotype (designated here): A) "Phelipaea flava / mihi (no. 94) / In arena mobili versus / mare caspium prope / Karawansarai [Karvansaray / Karabansarai] Eszek. Ad / Calligoni radices / d. 29 april: mens. 1830 / Enum. cauc. casp. No. 899" [manu C. A. Meyer]. B) "Cistanche tubulosa (Schenk) K.Wight / var. flava (C.A.Mey.) G.Beck / specim. Meyeri authenticum / 1947 11. X Determ. I. Novopokrovsky’ [handwritten by N. N. Tzvelev added ‘Typus!’]" (LE01064312, Supporting information; isolectotypes LE01064314, P02970920; authentic specimen LE01064313)» (Uhlich & Rätzel 2024: 22, Suppl. Fig. 32 ["Lectotype (P02970920, designated here)"] [the photo correspond to an isolectotype]).

1. "Phelipaea flava / mihi (Nş 94) / In arena mobili versus / mare Caspium prope Karavansarai [Karvansaray / Karabansarai] Eszek. Ad / ad Calligoni radices / d. 29 aprili: ... 1830 / Enum. cauc. casp. № 899 [leg. Meyer]". – 2. "Cistanche tubulosa (Schenk) R. Wight / var. flava (C.A. Mey.) G. Beck / Specim. Meyeri authenticum! Typus! / 1947.11.X  Determ. I. Novopokrovsky" (http://re.herbariumle.ru/01064312 - LE 01064312.jpg [Record creation: 2019-11-13, Ivan Tatanov, PhotoScan D1] - LE s.n. Meyer-PiwowarczykDSC09957, syntypes); 1. "Phelipaea flava / Enum. cauc. casp. № 899 / Herb. Fischer [leg. Meyer]". – 2. "Cistanche tubulosa Wight / Dr. G. de Beck [m. Beck]". – 3. "I. Novopokrovsky. Notae criticae. / Cistanche flava (C.A.M.) / Korsh. / Cotypus! / 1948.21.V" (http://re.herbariumle.ru/01064314 - LE 01064314.jpg [Record creation: 2019-11-13, Ivan Tatanov, PhotoScan D1] - LE s.n. Meyer-PiwowarczykDSC09956, syntype); 1. "Phelipea flava C.A.M. / ad Calligon. radei / Mar. Caspium". – 2. "Phelipea flava C. A. Meyer. / Mare caspium". – 3. "I. Novopokrovsky. Notae criticae. / Cistanche flava (C. A. Mey.) / Korsh. / Specim. Meyeri authenticum! / 1948.11.V". – 4. "stigma reniforme / thecae subotusae / [Stigma and anthera drawings]" (http://re.herbariumle.ru/01064313 - LE 01064313.jpg [Record creation: 2019-11-13, Ivan Tatanov, PhotoScan D1] - LE s.n. Meyer-PiwowarczykDSC09955, syntypes); "Herb. Acad. Petrop. / N. 899 / Phelipaea flava C.A. M. / Fl. Caucas. legit C.A. Meyer 1829. 1830. // Herb. Al. de Bunge" (https://science.mnhn.fr/institution/mnhn/collection/p/item/p02970920 - https://science.mnhn.fr/institution/mnhn/collection/p/item/p02970920 - P 02970920.jpg - hb. E. Cosson - https://www.gbif.org/occurrence/437278437 - https://explore.recolnat.org/occurrence/A380F88018A3442A8448B05FDD3A95AD; rev. Ó. Sánchez Pedraja 2016, syntype).

Designated by: "Type in Leningrad [LE]" (Novopokrovsky & Tzvelev 1958: 24[22], sub C. flava). "Azerbaijan, between Baku and Salijanami, Novopokrosky 899 (LE !, Type)" (Saeidi & Shahi 2008: 98). "Lectotype ... LE01064312, Supporting information; isolectotypes LE01064314, P02970920; authentic specimen LE01064313) by Uhlich & Rätzel in Nordic J. Bot. e04155: 22, Suppl. Fig. 32 ["Lectotype (P02970920, designated here)"] [the photo (f. 32) corresponds to an isolectotype] (2024).

Host: "Parasitic on species of Calligonum L. [Polygonaceae Juss.]" (Novopokrovsky & Tzvelev 1958: 24[22], sub C. flava). "Calligonum, Suaeda, Alhagi" (Saeidi & Shahi 2008: 97).

Comments:

Morphological observation:

"The bracts of C. tubulosa are oblate or ovate-lanceolate, while the bracts of C. flava are oblate or lanceolate-linear. Also the bractlets in C. tubulosa are smaller than the calyx, whereas they are as long as calyx or longer than the calyx in C. flava" (Saeidi & Shahi 2008: 96)

Taxonomic observations:

Cistanche christisonioides Beck in Engl., Pflanzenr. 96: 36 (1930) (Ataei 2017: 84). Ind. loc.: Pakistan: "Ost-Indien: Im Punjab [Pănjáb] bei Lahóre, im März 1857 (Schlagintweit n. 10485 im Herb. München [M], Wien  [W]!)" (Beck 1930: 36). Type: 1. "Herbarium Schlagintweit ... / Northwestern India / Province: Pănjáb / Locality: Lahóre ... / Collected 10 to 14 March 1857 / ... No. ... 10485" . – 2. "... / Cistanche christisonioides / m. / G. Beck [m. Beck]" (M 0185673 - https://www.gbif.org/es/occurrence/1098987633). Obs.: According to Ataei (2017: 84) it is a synonym of C. flava (C.A. Mey.) Korsh.

Genetic observations:

Chromosomatic number:

Ic.: a, Saeidi & Shahi 2008: 98 f. 1; b, Ataei 2017: 98 f. 4.12; b, Aldughayman & al. 2024: f. 3, sub C. tubulosa; c, photos: habitat, habit, scales, flower lateral view, flower-section, flower frontal view (https://www.plantarium.ru/page/image/id/807210.html by Александр Фатерыга [Alexander Fateryga], sub C. flava, in Plantarium; accessed Jun 2024)

Distribution: Afghanistan [n.v.], Azerbaijan, Iraq [in error for C. tubulosa], Iran [needs to be review], Kazakhstan, Pakistan [see C. christisonioides], Turkmenistan and Uzbekistan.

Distribution: "Caucasus: eastern Transcaucasia. Soviet Central Asia: Kara Kum [Karakum / Gara-Gum Desert], Kyzyl Kum, Balkhash Region [Balkhash District / Балхашский район / Балқаш ауданы] (southern part), Pamiro-Alai [Pamir-Alay / Памиро-Алай] (western part). General distribution: Iran" (Novopokrovsky & Tzvelev 1958: 24[22], sub C. flava). "Persia (?), Karakum, Pamiro-Alaj" (Schiman-Czeika 1964: 21, sub C. flava).

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Specimens examined and literature records.

Afghanistan /  افغانستان / Afġānistān / Afganistán [n.v.] (Ataei 2017: 85, sub C. flava)

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Azerbaijan / Azərbaycan / Азәрбајҹан / Azerbaiyán / Aserbaidschan / Azerbaďdjan / Азербайджан (Ledebour 1849: 314, sub P. flava; Beck 1930: 33, sub C. tubulosa var. flava; Novopokrovsky & Tzvelev 1958: 24[22], sub C. flava)

Baku Economic Region / Bakı iqtisadi rayonu / Бакинский экономический район: 1. "Phelipaea flava / mihi (Nş 94) / In arena mobili versus / mare Caspium prope Karavansarai [Karvansaray / Karabansarai] Eszek. Ad / ad Calligoni radices / d. 29 aprili: ... 1830 / Enum. cauc. casp. № 899 [leg. Meyer]". – 2. "Cistanche tubulosa (Schenk) R. Wight / var. flava (C.A. Mey.) G. Beck / Specim. Meyeri authenticum! Typus! / 1947.11.X  Determ. I. Novopokrovsky" (http://re.herbariumle.ru/01064312 - LE 01064312.jpg [Record creation: 2019-11-13, Ivan Tatanov, PhotoScan D1] - LE s.n. Meyer-PiwowarczykDSC09957, syntypes of P. flava); 1. "Phelipaea flava / Enum. cauc. casp. № 899 / Herb. Fischer [leg. Meyer]". – 2. "Cistanche tubulosa Wight / Dr. G. de Beck [m. Beck]". – 3. "I. Novopokrovsky. Notae criticae. / Cistanche flava (C. A. M.) / Korsh. / Cotypus! / 1948.21.V" (http://re.herbariumle.ru/01064314 - LE 01064314.jpg [Record creation: 2019-11-13, Ivan Tatanov, PhotoScan D1] - LE s.n. Meyer-PiwowarczykDSC09956, syntype of P. flava); 1. "Phelipea flava C.A.M. / ad Calligon. radei / Mar. Caspium". – 2. "Phelipea flava C. A. Meyer. / Mare caspium". – 3. "I. Novopokrovsky. Notae criticae. / Cistanche flava (C. A. Mey.) / Korsh. / Specim. Meyeri authenticum! / 1948.11.V". – 4. "stigma reniforme / thecae subotusae / [Stigma and anthera drawings]" (http://re.herbariumle.ru/01064313 - LE 01064313.jpg [Record creation: 2019-11-13, Ivan Tatanov, PhotoScan D1] - LE s.n. Meyer-PiwowarczykDSC09955, syntypes of P. flava); "Herb. Acad. Petrop. / N. 899 / Phelipaea flava C.A. M. / Fl. Caucas. legit C.A. Meyer 1829. 1830. // Herb. Al. de Bunge" (https://science.mnhn.fr/institution/mnhn/collection/p/item/p02970920 - https://science.mnhn.fr/institution/mnhn/collection/p/item/p02970920 - P 02970920.jpg - hb. E. Cosson - https://www.gbif.org/occurrence/437278437 - https://explore.recolnat.org/occurrence/A380F88018A3442A8448B05FDD3A95AD; rev. Ó. Sánchez Pedraja 2016, syntype of P. flava); "An den Küsten des Caspischen Meeres im Dünensande bei Erzek zwischen Baku und Sallian (C. A. Meyer)" (Beck 1930: 33, sub C. tubulosa var. flava); "Described from Azerbaidzhan (between Baku and Salyans). Type in Leningrad. " (Novopokrovsky & Tzvelev 1958: 24[22], sub C. flava).

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Iraq / Irak / Ирак (Ataei 2017: 87, sub C. flava; Aldughayman & al. 2024: f. 3, sub C. tubulosa)

Diyala Governorate: 1. "W847 Iraq / Cistanche / Shahraban [Miqdadiyah] 11 Apr.57 / Occasional on low sand / dunes, parasite on ? Ephedra / enclosed. / Yellow throat, purple outside / tube and purple lobes. / ... / host abundant on tops of dunes & on sides, parasites only on sides...". – 2. "C. tubulosa ...". – 3. "Cistanche flava (C.A. Mey.) Korsh. N. Ataei 2012.02.03" (http://specimens.kew.org/herbarium/K001222533 - https://d2seqvvyy3b8p2.cloudfront.net/e813d90d35e6404d7ebbcb8bfa39eb6d.jpg - https://data.kew.org/images...8cba4ddd-bcd9-4789-a149-f91c9e8a1817 (Aldughayman & al. 2024, sub C. tubulosa)); 1. idem . – 2. "C. tubulosa (Schenk) R. Wight / 1972 det. A. Gilli". – 3. "Cistanche flava (C.A. Mey.) Korsh. subsp. flava / N. Ataei Mar 2012" (https://data.rbge.org.uk/herb/E00029723 - E00029723.zip - https://www.gbif.org/occurrence/575243924; rev. Ó. Sánchez Pedraja Jun 2024 (Aldughayman & al. 2024, sub C. tubulosa)).

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Iran / Irán / Иран / Persia / Persien [needs to be review] (Beck 1930: 33, sub C. tubulosa var. flava; Novopokrovsky & Tzvelev 1958: 24[22], sub C. flava; Schiman-Czeika 1964: 21, "Persia (?)", sub C. flava; Saeidi & Shahi 2008: 98, sub C. flava subsp. flava; Saeidi & Shahi 2008: 99, sub C. flava subsp. bicolor; Ataei 2017: 86-88, sub C. flava; Ataei 2017: 87, loc. class. of C. flava subsp. brevibracteata)

Bushehr Province: 1. " Davis & Bokhari (D.55976) / Cistanche tubulosa (Schenk) Wight / Iran. D6 (Times Atlas): Mond river / N of Kaki. 100 m. Lowe sand / dunes on S. bank. On Haloxylon. / Flowers yellow (bud pale violet) / 27 March 1974". – 2. "Cistanche flava (C.A. Mey.) Korsh. subsp. brevibracteata / subsp. nov. / Det.: Najibeh Ataei May 2014 / ..." (https://data.rbge.org.uk/herb/E00028866 - E 00028866.zip - https://www.gbif.org/occurrence/575300308 (Ataei 2017: 88, sub C. flava subsp. brevibracteata); rev. Ó. Sánchez Pedraja Jun 2024, bracteoles oblong-lanceolate shorter than calyx with scarious margins, corolla lobes tinged purple, anthers obtuse, see C. tubulosa (Schenk) Hook. f.)).

Isfahan Province: "Persien! bei Ispahan (Aucher n. 5090)" (Beck 1930: 33, sub C. tubulosa var. flava); "K.H. et F. Rechinger, Iter iranicum II, 1948 / ... / No. 2704 / Cistanche tubulosa (Schenk) R. Wight / Prov. Semnan et Yezd: / Ad margines deserti inter Nain [Isfahan] et Aghda [Aqda (Yazd)], / ca. 1000 m. / 21.IV det. H. Schiman-Czeika, 1963". – 2. " Cistanche flava (C. A. Mey.) Korsh. subsp. flava / det. Najibeh Ataei" (https://herbarium.bgbm.org/object/B100264803 - B 10 0264803 - https://image.bgbm.org/images/internal/HerbarThumbs/B100264803_1700 - https://www.gbif.org/fr/occurrence/911748652; rev. Ó. Sánchez Pedraja Jun 2024, cf. [specimen with lanceolate scales and ovate-lanceolate bracts, but bad preserved] (Ataei 2017: 85, sub C. flava subsp. flava).

Kerman Province: 1. "J. Bornmüller. Iter Persico-turcicum / 1892-93 / 3900 / Ph. flava C. A. Mey. / Boiss. fl. Or. / (Phelipaea) Cistanche tubulosa Wight / f. flava (C.A. Mey.)  sec. Dr. G. v Beck / ... / Persia austro-or.: in desertis ad Kebuter Khan [Kabutar Khan / Kabootarkhan (Rafsanjan County)] / inter Yesd [Yazd / Yezd, Yazd County] et Kerman) / 16-1800 m.s.m. / 1892.18.IV legit: J. Bornmüller". – 2. "nr 3900 Cistanche tubulosa Wight / f. flava (C.A. Mey.) [m. Beck?]". – 3. "nr 3900 ex G. Beck l.c.p. 33 / Cistanche tubulosa Wight / α. flava (C.A. Mey.) [m. Beck?]" (https://herbarium.bgbm.org/object/B100264810 - B 10 0264810 - https://www.gbif.org/fr/occurrence/144848592; rev. Ó. Sánchez Pedraja May 2018, specimens with long scales and bracts, but bad preserved (Bornmüller 1904: 686, sub C. tubulosa (Schenk) R. Wight)); "... / No. 3900 / Ph. flava C.A.M. / Boiss. Or. / (Phelipaea) Cistanche tubulosa Wight / f. flava (C.A. Mey.)  sec. Dr. G. v Beck / Persia austro-or.: in desertis ad / Kebuter Khan [Kabutar Khan / Kabootarkhan (Rafsanjan County)] / (inter Yesd [Yazd / Yezd, Yazd County] et Kerman) / 16-1800 m.s.m. / 1892.18.IV legit: J. Bornmüller" (https://herbarium.bgbm.org/object/B100264811 - B 10 0264811 - https://image.bgbm.org/images/internal/HerbarThumbs/B100264811_1700 - https://www.gbif.org/fr/occurrence/144848593; rev. Ó. Sánchez Pedraja Jun 2024 [specimens with ovate-lanceolate scales and bracts, but bad preserved] (Bornmüller 1904: 686, sub C. tubulosa (Schenk) R. Wight)); "Persia orientalis: in desertis inter Yesd [Yazd / Yezd, Yazd County] el Kennan [Kermān / Kirman / Carmania, Kerman County] prope pagum Hormusabad, 16-1700 m. s. m.; 17.IV.1892; plantse nutrientes: Tamarix (Bornm., iter Persico-turcicum, n. 3899). — Die üppigen, bis 70 cm. hohen Exemplare sind an den obern Stengelteilen und Bracteen häufig mehr oder minder amethystfarbig, Blülhen goldgelb (Bornm., n. 3899b)." (Bornmüller 1904: 686, sub C. tubulosa (Schenk) R. Wight); "in Wüsten bei Khebuter Chan zwischen Yesd [Yazd / Yezd, Yazd County] und Kerman bei 1600—1800 m im April (Bornmüller, Iter pers.-turc. 1892—1893, n. 3900!)" (Beck 1930: 33, sub C. tubulosa var. flava); 1. "Gabriel: Iter Irano-gedrosiacum / a. 1937 / No. 169 / Cistanche tubulosa (Schenk) Wight / ... / Kŭh Čah-Rŭi [Kuh-i Chah Rui (Lut Desert / Dasht-e Lut)] ...1937-03-15, Leg.: Dr. Alfons et Agnes Gabriel / 15-18,III.1937". – 2. " Cistanche flava (C. A. Mey.) Korsh. subsp. flava / det. Najibeh Ataei" (https://www.jacq.org/detail.php?ID=528493 - http://herbarium.bgbm.org/object/B100264807 - https://www.gbif.org/fr/occurrence/911748646; rev. Ó. Sánchez Pedraja Jun 2024, cf. [specimen with lanceolate scales and ovate-lanceolate bracts, but bad preserved] (Schiman-Czeika 1964: 21, sub C. tubulosa (Schenk) R. Wight)); 1. "K.H. Rechinger, Iter Orientale 1965 / Flora of Iran / Cistanche tubulosa (Schenk) R. Wight / Prov. Kerman: 10 km E. of Nasratabad Sipi / 27th March, 1965 / Coll. Jennifer Lamond No. 111". – 2. "1966 rev. A. Gilli". – 3. "Cistanche flava (C.A. Mey.) Korsh. subsp. flava / Det.: Najibeh Ataei May 2014 / ..." (https://data.rbge.org.uk/herb/E00028868 - E 00028868.zip - https://www.gbif.org/occurrence/575351331 (Ataei 2017: 87, sub C. flava); rev. Ó. Sánchez Pedraja Jun 2024, sub C. tubulosa (Schenk) Hook. f. [scales and bracts short, ovate-lanceolate]); 1. "K.H. Rechinger, Iter Orientale 1965 / Flora of Iran / Cistanche tubulosa (Schenk) R. Wight / Prov. Kerman: 10-20 km E. of Nasratabad Sipi / Growing on Ephedra. Fls. lemon / 27th March, 1965 / Coll. Jennifer Lamond No. 115". – 2. "1966 rev. A. Gilli". – 3. "Cistanche flava (C.A. Mey.) Korsh. subsp. flava / Det.: Najibeh Ataei May 2014 / ..." (https://data.rbge.org.uk/herb/E00028869 - E00028869.zip  - https://www.gbif.org/fr/occurrence/575098663 (Ataei 2017: 87, sub C. flava); rev. Ó. Sánchez Pedraja Jun 2024, sub C. tubulosa (Schenk) Hook. f. [scales and bracts long, lanceolate to ovate-lanceolate, flowers with tigned lobes]).

Semnan Province: "K.H. Rechinger, Iter iranicum IX, 1975 / 50123 / Cistanche laxiflora Aitch. et Hemsl. / Persia: C: Kavir (Kavir protected region [Kavir National Park]): / Siah Kuh, In ditione refugii Karvan-Sarai Shah Abbas / 34°44′00''N, 52°10′00''E, 1100-1300 m / 23.V.1975" (https://www.jacq.org/detail.php?ID=529264 - https://herbarium.bgbm.org/object/B100264822 - B 10 0264822.jpg - https://www.gbif.org/es/occurrence/144848604 (Ataei 2017: 87, sub C. flava)); idem (W (Ataei 2017: 87, sub C. flava)); "Cistanche tubulosa (Schenk) R. Wight / Touran Protected Area [Khar Turan National Park, Semnan prov.] / SE of Shahrud at road / to 1,5 km SW of Chejam / sand fields, Haloxylon persicum ass. / Alt.: 1100 m. / 23.04.1978 / Coll. H. Freitag 14695" (https://herbarium.bgbm.org/object/B100264808 - https://image.bgbm.org/images/internal/HerbarThumbs/B100264808_1700 - https://www.gbif.org/occurrence/911748649 (Ataei 2017: 88, sub C. flava subsp. brevibracteata); rev. Ó. Sánchez Pedraja Jun 2024, bracteoles oblong-lanceolate shorter than calyx, corolla lobes tinged purple, anthers obtuse, see C. tubulosa (Schenk) Hook. f.)); "Shahrud, to Ahmad abad, Iranshahr s.n.; Sharud, Kalat Hizomi, Shahi & Saeidi 13230" (Saeidi & Shahi 2008: 98, "hairy along edges" [?], sub C. flava subsp. flava); "protected region of Turan [Khar Turan National Park], 3 km Chahjam to Turud, 1100 m, Freitag & Mozaffarian 28437" (Saeidi & Shahi 2008: 99, sub C. flava subsp. bicolor).

Southern Khorassan Province: "Herbar. Bungeanum. / Iter persicum. / pr. Sser-tschah [Sar Chah-e Tazian / Sar-i-tschah / Sar-I-Chah, 33°01'07.0"N 59°33'38.0"E 33.018611, 59.560556] Persiae mediae orientalis. Mart. 1859." (http://coldb.mnhn.fr/catalognumber/mnhn/p/p02983449 - P 02983449.jpg - hb. E. Cosson - http://sonneratphoto.mnhn.fr/2011/10/14/3/P02983449.jpg - http://www.gbif.org/occurrence/438437918 - https://explore.recolnat.org/occurrence/4F375F34F1154444A3A047B0730A9BE3; rev. Ó. Sánchez Pedraja Feb 2015 [stem with long narrow lanceolate scales and flowers with ovate-lanceolate bracts, see C. flava]); "bei Ssertschah [Sar Chah-e Tazian / Sar-i-tschah / Sar-I-Chah, 33°01'07.0"N 59°33'38.0"E 33.018611, 59.560556im südöstlichen Persien [Persiae mediae orientalis Mart. 1859 (see LE 00010740)] (Bunge)" (Beck 1930: 33, sub C. tubulosa var. flava); "Type: Iran. Southern Khorassan: 40-42 km after Nehbandan towards Zabol, 31°29'29.0"N 60°36'34.0"E 31.491389, 60.609444, 807-825 m, 30 March 2011, Akhani, Noormohammadi & Samadi 22074 (holotype TUH)" (Ataei 2017: 87, holotype of C. flava subsp. brevibracteata)).

Yazd Province: "K.H. et F. Rechinger, Iter iranicum II, 1948 / ... / No. 2704 / Cistanche tubulosa (Schenk) R. Wight / Prov. Semnan et Yezd: / Ad margines deserti inter Nain [Isfahan] et Aghda [Aqda (Yazd)], / ca. 1000 m. / 21.IV det. H. Schiman-Czeika, 1963". – 2. " Cistanche flava (C. A. Mey.) Korsh. subsp. flava / det. Najibeh Ataei" (https://herbarium.bgbm.org/object/B100264803 - B 10 0264803 - https://image.bgbm.org/images/internal/HerbarThumbs/B100264803_1700 - https://www.gbif.org/fr/occurrence/911748652; rev. Ó. Sánchez Pedraja Jun 2024, cf. [specimen with lanceolate scales and ovate-lanceolate bracts, but bad preserved] (Ataei 2017: 85, sub C. flava subsp. flava).

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Kazakhstan / Қазақстан / Qazaqstan / Казахстан / Kazajistán (Novopokrovsky & Tzvelev 1958: 24[22], sub C. flava; Aldughayman & al. 2024: f. 4C, D, sub C. flava; Fateryga 2024, sub C. flava)

Almaty Region / Almaty Oblysy / Алматы облысы / Алматинская область: "Balkhash Region (southern part) [Balkhash District / Балхашский район / Балқаш ауданы]" (Novopokrovsky & Tzvelev 1958: 24[22], sub C. flava); "Цветущее растение. Казахстан, Алматинская обл., Государственный национальный парк "Алтын Эмель" [Altyn-Emel National Park], возле "Поющего бархана [Singing Dune]". 23.04.2016 (https://www.plantarium.ru/page/image/id/450419.html, https://www.plantarium.ru/page/image/id/450420.html and https://www.plantarium.ru/page/image/id/450421.html by Лина Вальдшмит [Lina Valdshmit], таксон определил(а) Владислав Григоренко [Vladislav Grigorenko], sub C. flava, in Plantarium; accessed Jun 2024 [stem with long narrow lanceolate scales and flowers yellowish tinged with purple and glabrous] (Aldughayman & al. 2024: f. 4C, D, sub C. flava)); "Цветущие растения. Казахстан, Алматинская обл. [Almaty Region], нац. парк "Алтын-Эмель" [Altyn-Emel National Park], подножие Поющего бархана [Singing Dune], слабозакреплённые пески. 3 июня 2017 г." (https://www.plantarium.ru/page/image/id/669191.html by Владимир Панкратов [Vladimir Pankratov], таксон определил(а) Александр Фатерыга [Alexander Fateryga], sub C. flava, in Plantarium; accessed Jun 2024).

Jetisu Region / Zhetysu Region / Жетісу облысы / Жетысуская область: Singing Dune (Altyn-Emel National Park), May 2021 (https://maps.app.goo.gl/TFKKKnx3bGhpyR3m7 by 홍명호 [Hong Myung-ho]; accessed et rev. Ó. Sánchez Pedraja Jun 2024, on ?/cf. Calligonum sp. [stem with long narrow lanceolate scales and flowers pale yellow, slightly dyed purple, glabrous and with long bracts similar to the scales]); "Цветок. Казахстан, Жетысуская обл. [Jetisu Region], нац. парк "Алтын-Эмель" [Altyn-Emel National Park], Айгайкум (Поющий бархан) [Aigaykum (Singing Dune), 43°52'47.6"N 78°33'50.4"E 43.879889, 78.564000], дюны. 4 мая 2024 г." (https://www.plantarium.ru/page/image/id/807210.html by Александр Фатерыга [Alexander Fateryga], sub C. flava, in Plantarium; accessed Jun 2024 [very illustrative images]); "Кербулакский район [43°52'41.2"N 78°33'36.8"E 43.878120, 78.560210, Singing Dune (Altyn-Emel National Park)], 4 May 2024" (https://www.inaturalist.org/observations/214581475 - https://www.gbif.org/occurrence/4863800934 by https://www.inaturalist.org/people/mylabris, det. vladimir_epiktetov, sub C. flava, in iNaturalist; accessed Jun 2024); "Кербулакский район [Kerbulak District, 43°52'36.9"N 78°33'40.6"E 43.876917, 78.561278, Singing Dune (Altyn-Emel National Park)], 5 May 2024" ( https://www.inaturalist.org/observations/213719950 - https://www.gbif.org/occurrence/4855556571 by https://www.inaturalist.org/people/madkiba, sub C. salsa, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Apr 2015, sub C. flava).

Qyzylorda Region / Қызылорда облысы / Кызылординская область [formerly Kyzyl-Orda Region]: "Kyzyl Kum" (Novopokrovsky & Tzvelev 1958: 24[22], sub C. flava).

Unknown location: "Цветущее растение в пустыне. Казахстан. Середина мая." (http://www.plantarium.ru/page/image/id/59887.html by Рустам Бобров [Rustam Bobrov], 2010; растение определил(а) [det.] Андрей Ковальчук [Andriy Kovalchuk], sub Cistanche, in Plantarium; accessed et rev. Ó. Sánchez Pedraja Apr 2015 et Jun 2024 [cf. C. flava]).

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Pakistan / Pakistán / Paquistán / Пакистан (Beck 1930: 36, loc. class. of C. christisonioides; Ataei 2017: 84, type of C. christisonioides; Ataei 2017: 87, sub C. flava subsp. flava)

Balochistan Pakistan: 1. "Date May 1889 / Altd. 6000 feet / Locality Surkhab [Pishin District] / Cistanche / Field No. 3387 / Herb. J.H. Lace". – 2. "Cistanche flava  (C.A. Mey.) Korsh. subsp. flava / Det.: Najibeh Ataei May 2014 / ..." (https://data.rbge.org.uk/herb/E00028906 - E 00028906.zip - https://www.gbif.org/occurrence/574907691 (Ataei 2017: 87, sub C. flava subsp. flava); rev. Ó. Sánchez Pedraja Jun 2024 [stem with long narrow lanceolate scales and flowers with long bracts similar to the scales, cf. C. flava]).

Punjab Province: 1. "Herbarium Schlagintweit ... / Northwestern India / Province: Pănjáb / Locality: Lahóre ... / Collected 10 to 14 March 1857 / ... No. ... 10485" . 2. "... / Cistanche christisonioides / m. / G. Beck [m. Beck]" (M 0185673 - https://www.gbif.org/es/occurrence/1098987633, type of C. christisonioides); "Ost-Indien [Pakistan]: Im Punjab [Pănjáb] bei Lahóre, im März 1857 (Schlagintweit n. 10485 im Herb. München, Wien  [W]!)" (Beck 1930: 36, loc. class. of C. christisonioides).

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Turkmenistan / Türkmenistan / Turkmenia / Туркмения / Turkmenien / Turcomannia (Karelin 1839: 165, loc. class. of P. gigantea; Ledebour 1849: 314, sub P. flava; Beck 1930: 37, sub C. flava; Novopokrovsky & Tzvelev 1958: 24[22], sub C. flava)

Ahal Region / Ahal welaýaty / Ахалский велаят: "Kara Kum" (Novopokrovsky & Tzvelev 1958: 24[22], sub C. flava); "Цветущие растения. Туркменистан, Центр. Каракумы [Karakum / Gara-Gum Desert], заповедник "Берекетли Каракум" [Bereketli Garagum State Reserve], незакр. пески. 11.04.2019" (https://www.plantarium.ru/page/image/id/610375.html by Александр Павленко [Alexandr Pavlenko], sub C. flava, in Plantarium; accessed Jun 2024 [stem with long narrow lanceolate scales and flowers yellowish-white tinged with purple]).

Balkan Region / Balkan welaýaty / Балкан велаяты / Балканский велаят: "In insula Ogurtsch'nsk [Ogurtschinsky Island / Ogurchinsky / Ogurja Ada / Aidak Island in the sutheastern part of Caspian Sea]" (Karelin 1839: 165, loc. class. of P. gigantea). "Siberia uralensi in insula Ogurtschik [Ogurtschinsky Island / Ogurchinsky / Ogurja Ada / Aidak Island] m. caspii [Caspian Sea]! (Karel)" (Ledebour 1849: 314, sub P. flava). "auf der Insel Ogurtschinsk am Caspi See (Karelin als Ph. gigantea!)" (Beck 1930: 37, sub C. salsa).

Lebap Region / Lebap welaýaty / Лебап велаяты / Лебапский велаят: "... 6499. Cistanche flava (C.A. Mey.) Korsh. / ... / ... — Turcomania, / in arenis mobillibus prope st. v. f. Repetek [Репетек, near Türkmenabat]. Leg. A. Vvedensky, det. / Z. Filimonova. / 1927.IV.25" (https://science.mnhn.fr/institution/mnhn/collection/p/item/p02983475 - P 02983475.jpg - ex hb. Prievidza, Slovakia (HMP) - http://www.gbif.org/occurrence/439524382 - https://explore.recolnat.org/occurrence/421EC6195F194A35997093641E367643; rev. Ó. Sánchez Pedraja Sep 2016); idem (https://plant.depo.msu.ru/open/public/en/item/MW0877616 - MW 0877616.jpg; rev. Ó. Sánchez Pedraja May 2018); idem (https://plant.depo.msu.ru/open/public/item/MW0877615 - MW 0877615.jpg; rev. Ó. Sánchez Pedraja May 2018); idem (PE 01542440 - https://www.cvh.ac.cn/spms/detail.php?id=eb93d720; rev. Ó. Sánchez Pedraja May 2018); idem (LE s.n. - PiwowarczykDSC08237); idem (LE s.n. - PiwowarczykDSC08241); idem (https://data.rbge.org.uk/herb/E00029684 - E 00029684.zip - https://www.gbif.org/fr/occurrence/575336645; rev. Ó. Sánchez Pedraja Jun 2024); 1. idem. –  2. "... DNA ... / Najibeh Ataei ... / Determinavit 29.07.2010" (http://www.botanicalcollections.be/specimen/BR0000008058218 - BR 0000008058218.jpg - https://www.gbif.org/es/occurrence/4072423205 (Ataei 2017: 87, sub C. flava)); "Репетексая песчаная станция /... 38°35'00.0"N 63°10'00.0"E 38.583333, 63.166667 ... / Cistanche flava (C.A. Mey.) Korsh. / Sandy desert Karakum [Каракум] Barchan [Бархан] / 3 km NE of ... / Barkhan-sands [Barchan, Repetek Station / Репетекский государственный биосферный заповедник] / 3.V.1935 / Е. Благовещенский [E. Blagoveschenskiy] / Repetek Sand Station" (http://n2t.net/ark:/65665/m3362f8bd8-a30e-47f2-9667-f75dab4650e3 - http://n2t.net/ark:/65665/399f22de5-f0a7-45b3-b758-c04b935edf72 - US 1665968 - NMNH-03926281.jpg - https://www.gbif.org/fr/occurrence/2625849768; rev. Ó. Sánchez Pedraja Feb 2024 [illustrative specimen]); "Цветущее растение на корнях Artemisia dimoana [?, see Calligonum]. Туркменистан, ЮВ Каракумы [SE Karakum / Gara-Gum Desert], Репетекский заповедник [Repetek Biosphere State Reserve], слабозакр. пески. 04.04.2019" (https://www.plantarium.ru/page/image/id/609731.html by Александр Павленко [Alexandr Pavlenko], sub C. flava, in Plantarium; accessed Jun 2024; "Цветущие растения. Туркменистан, ЮВ Каракумы [SE Karakum / Gara-Gum Desert], Репетекский заповедник [Repetek Biosphere State Reserve], незакр. пески. 05.04.2019" (https://www.plantarium.ru/page/image/id/610374.html by Александр Павленко [Alexandr Pavlenko], sub C. flava, in Plantarium; accessed Jun 2024); "Lebap [38°32'19.2"N 63°23'14.0"E 38.538660, 63.387220 (Repetek Biosphere State Reserve)], 5 May 2024" (https://www.inaturalist.org/observations/218476546 by https://www.inaturalist.org/people/markltask [Mark Tasker], sub C. tubulosa, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Jun 2024, C. flava, long narrow lanceolate bracts).

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Uzbekistan / Uzbekistán / Ouzbékistan / Usbekistan / Узбекистан (former Uzbek SSR / Ўзбекистон ССР / Узбекская ССР) (Beck 1930: 33, sub C. tubulosa; Novopokrovsky & Tzvelev 1958: 24[22], sub C. flava; Ataei 2017: 87, sub C. flava; Ataei 2017: 92, syntype of of C. mongolica [= C. laxiflora subsp. laxiflora])

Andijan Region / Andijon viloyati / Андижон вилояти / Андижанская область: 1. "P. ..., 1906 / N.ş 229 Cistanche salsa / Chirabad [Chinobod / Chinabad / Чинобод / Чинабад]: Sables Patta-Koum [Kyzyl Kum desert, Bukhara Region (Emirate of Bukhara)]. / 11 avril  Roshewitz, Plantae bucharicae 1906.". – 2. "Cistanche flava" (http://coldb.mnhn.fr/catalognumber/mnhn/p/p02983457 - P 02983457.jpg - http://www.gbif.org/occurrence/438609505 - https://explore.recolnat.org/occurrence/966EC3AFA9CB4C28A6E21A6DB111C248; rev. Ó. Sánchez Pedraja Mar 2016, glabrous!, it does not seem to be C. salsa (Ataei 2017: 87, sub C. flava)); "Turkmenien: Buchara [Bukhara / Buxoro / Бухараa]! auf dem Baba-Tag (Roshewitz, pl. bucharicae [1906] n. 752! in Herb. Berlin) ... Turkestan: zwischen den Brunnen Jeschak-rabatt und Rofatak sowie zwischen Tschardschui [Khanate of Bokhara] uud dem Brunnen Salim, im Mai fruchtend (Regel)" (Beck 1930: 33, sub C. tubulosa (Schenk) R. Wight); "Buchara!: bei Patta Kum [Kyzyl Kum desert, Bukhara Region (Emirate of Bukhara)] im Sande, April 1906 (Roshewitz, PI. Buchar. (1906) n. 229! in Herb. Berlin)" (Beck 1930: 34, loc. class. of C. mongolica Beck); "Uzbekistan. Buchara: Patta Kum im Sande, 1906, Roshewitz Pl. Buchar n. 229 (B)" (Ataei 2017: 92, syntype of C. mongolica [= C. laxiflora subsp. laxiflora]).

Bukhara Region / Buxoro viloyati / Бухоро вилояти / Бухарская область: "Cistanche tubulosa Wight / Tengi Charam [Tengi Charam, montagne de Baissoun (Boukharie)], 23 avril / M. Capus. Plantes du Turkestan. 1881 / Reçu le 27 Octobre 1881" (http://coldb.mnhn.fr/catalognumber/mnhn/p/p02983453 - P 02983453.jpg - http://sonneratphoto.mnhn.fr/2011/10/14/3/P02983453.jpg - http://www.gbif.org/occurrence/438437917 - https://explore.recolnat.org/occurrence/B83C68237CA64AD5AE6919A71F5BA093; rev. Ó. Sánchez Pedraja Feb 2015, stem with long narrow lanceolate scales and flowers with long bracts similar to the scales, cf. C. flava); 1. "5592. Cistanche flava (C.A. M.) / Бухарский Владислав [Bukhara Vladislav] ... Бархани [Bukhara] ... Calligonum / 1913.IV.18". – 3. "... 5592. / I. Novopokrovsky. Notae criticae. / Cistanche flava (C. A. Mey.) / Korsh. / ... 1947.17.V" (PE 01542441 - https://www.cvh.ac.cn/spms/detail.php?id=eb93d60e; rev. Ó. Sánchez Pedraja May 2018); "Romitan District [40°43'19.1"N 62°29'00.5"E 40.721970, 62.483470, near Qzilravat], 15 Apr 2013" (https://www.inaturalist.org/observations/69874865 - https://www.gbif.org/occurrence/3327710406 by https://www.inaturalist.org/people/lougarou, sub C. salsa, in iNaturalist, accessed et rev. Ó. Sánchez Pedraja Jul 2024, sub C. flava); "Romitan District [40°43'17.8"N 62°29'02.7"E 40.721610, 62.484090, near Qzilravat], 15 Apr 2013" (https://www.inaturalist.org/observations/69874956 - https://www.gbif.org/occurrence/3328024391 by https://www.inaturalist.org/people/lougarou, sub C. salsa, in iNaturalist, accessed et rev. Ó. Sánchez Pedraja Jul 2024, sub C. cf. flava [glabrous specimens, see https://www.inaturalist.org/observations/69874865]).

Navoiy Region / Навоий вилояти / Navoiy viloyati / Навоийская область: 1. "Cistanche flava!! / 8.VII.35 / Архиреев". – 2. "1935 Кызыл-Кум [Кызылкумы / Kyzylkum / Qizilqum] /  А.Д. Архиреев [A.D. Arkhireev]" (https://plant.depo.msu.ru/open/public/item/MW0877623 - MW 0877623.jpg; rev. Ó. Sánchez Pedraja May 2018); "Kyzyl Kum [Кызылкумы / Kyzylkum / Qizilqum]" (Novopokrovsky & Tzvelev 1958: 24[22], sub C. flava).

Qashqadaryo Region / Qashqadaryo viloyati / Қашқадарё вилояти / Кашкадарьинская область: "Pamiro-Alai [Pamir-Alay / Памиро-Алай] (western part)" (Novopokrovsky & Tzvelev 1958: 24[22], sub C. flava).

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