Accepted Name - Basionymum / Basionym - Synonyms - Ind. loc. [Indicatio locotypica (type locality)] - Lectotypus [Lectotype] - Designated by - Host - Comments - Chromosomatic number - Ic. [Images] - Distribution - Specimens Examined - References - Authors  - Cite us

ISSN: 2386-9666 - Last Updated: 1 May 2016


Accepted Name:

Orobanche densiflora Salzm. ex Bertol., Fl. Ital. 6: 444 (1846) ["O. densiflora Salzm., Pl. sicc. Hisp. Tingint. ex Reich. / O. crinita Reich., Cent. 7 p. 43. tab. 687 fig. 922. optim."] (cf. Reuter, 1847: 19; Cosson, 1849-1852: 42; Domina & Soldano, 2009; Domina & Raab-Straube, 2010)

Synonyms:

Homotypic synonyms:

Orobanche densiflora Salzm., Pl. Hisp. Tingit. Exsicc. (1825) [in sched., unpublished name] (cf. Beck, 1890: 232)

Orobanche densiflora Salzm. ex Rchb., Iconogr. Bot. Pl. Crit. 7: 43 (1829), pro syn. (cf. Reichenbach, 1829: 43, "O. densiflora Salzm. pl. hisp. tingit. exsicc.")

Orobanche densiflora Salzm. ex Mutel, Fl. Franç. 2: 351 (1835), pro syn. [nom. illeg.]. Obs. Orobanche densiflora sensu Mutel, Fl. Franç. 2: 351 (1835), non Orobanche densiflora Salzm. ex Mutel, Fl. Franç. 2: 351 (1835) [= Orobanche crinita Viv., Fl. Cors. Prodr.: 11 (1824)]

Orobanche densiflora Salzm. ex Steud., Nomencl. Bot. ed. 2: 230 (1841), pro syn.

Orobanche densiflora Salzm. ex Vis., Fl. Dalmat. 2: 180 (1847 [10-13 Nov. 1847]), nom. superfl. [nom. illeg.]

Orobanche densiflora Salzm. ex Reut. in DC., Prodr. 11: 19 (1847 [25 Nov. 1847]), nom. superfl. [nom. illeg.] (cf. Reuter, 1847: 19, "Salzm.! pl. tingit. exsicc. ... O. crinita Rchb. pl. crit. 7 fig. 922, non Viv !")

Orobanche densiflora Salzm. ex  Willk. in Willk. & Lange. Prodr. Fl. Hispan. 2(3): 622 (1870), nom. superfl. [nom. illeg.] (cf. Willkomm, 1870: 622, "Salzm. pl. Tingit. exs. ... O. crinita Rchb. pl. crit. VII, f. 922, nec Viv. teste Reut.")

Orobanche densiflora Salzm. ex Beck, Biblioth. Bot. 19: 232 (1890), nom. superfl. [nom. illeg.]

Orobanche densiflora var. typica J.A. Guim. in Brotéria 3: 143 (1904) ["α. typica O. densiflora Salzmann"], nom. illeg. (Art. 24.3)

Orobanche densiflora var. genuina Beck in Engl., Pflanzenr. 96: 194 (1930), nom. illeg. (Art. 24.3)

Orobanche densiflora var. genuina f. typica (J.A. Guim.) Beck in Engl., Pflanzenr. 96: 194 (1930), nom. illeg. (Art. 24.3)

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Heterotypic synonyms:

? = Orobanche densiflora var. macraixma J.A. Guim. in Brotéria 3: 144[-148] (1904). Syn.: Orobanche densiflora var. genuina f. macraixma (J.A. Guim.) Beck in Engl., Pflanzenr. 96: 195 (1930) (sec. M.J.Y. Foley in Flora iberica 14: 230. 2001b). Ind. loc.: [Portugal] "Centro littoral: Arredores de Lisboa: Tapada da Ajuda (parasita da Carlina corymbosa L., Welwitsch! 1846). / Alemtejo littoral: Collina da margem esquerda do Tejo (parasita da Thrincia hispida Rth., Daveau!).". Obs. The host-plants which have been indicated by the collectors do not seem to correspond to those of this species. In addition, we could see in the Paris herbarium a sheet from "Serpa [Beja District, Alentejo Region]" which had been collected and determined by Daveau as O. densiflora Salzm (P 04419810 [no. 20, Herbarium lusitanicum, Mai 1832]) but the specimen which contains do not correspond to this species, it would be O. rapum-genistae

? = Orobanche densiflora var. melitensis Beck in Engl., Pflanzenr. 96: 195 (1930) ["Var. β"]. Ind. loc: "Malta: an der Küste bei Marfa, im Mai (Sommier!)". Ic.: photos by S. Mifsud (accessed, 20 Nov. 2011). Herbarium Specimens: "Lectotypus: FI / a) O. densiflora Salzm. f. melitensis m. / differt a typo dentibus calycis partium alte connatis, corolla subcopiose glandulosa, glabrescens. / Ein kleines Exemplar behalten! Sehr interessanter Standort! [ho guardato un piccolo esemplare! Località molto interessante!] b) Stéphen Sommier – Plantae Melitenses / Orobanche densiflora Salzm. f. melitensis m. Beck / Insula Melita (hodie Malta) / Marfa – in litore arenoso. / 4 Maii 1907 legi / ex albo-flavescens / 55" (cf. Domina & Soldano, 2009). "Lo studio del materiale originale, conservato a Firenze (FI!), ha permesso di riferire questo taxon a O. sanguinea C. Presl." (cf. Domina & Soldano, 2009); "a) O. densiflora Salzm. / f. melitensis m. / differt a typo dentibus calycis / partium alte connatis, corolla subcopiose / glandulosa, glabrescens. / Ein kleines Exemplar / behalten! / Sehr interessanter Standort!" // b) Stéphen Sommier – Plantae Melitenses / Orobanche densiflora Salzm. / f. melitensis m. / Beck / Insula Melita (hodie Malta) / Marfa – in litore arenoso. / 4 Maii 1907 legi / ex albo-flavescens / 55 // c) Holotypus" (FI 002514).

= Orobanche crinita var. occidentalis M.J.Y. Foley in Anales Jard. Bot. Madrid 58: 226 (2001) [saltem p.p.]. Syn.: = Orobanche densiflora Salzm. ex Reut. (1847) [red flowering form with orangish-brown stigma]. Ind. loc.: "Holotype: Spain, Cádiz, Zahara de los Atunes, 26/27-IV-1993, Foley 243 (E [E 00053427]) /  Habitat: Maritime sands ... Parasitic upon Medicago spp.". Obs. some of the "Specimens examined" by Foley (2001a: 226) seem to correspond to stunted forms of O. foetida (cf. Foley, 2001a: 227; Pujadas & al., 2003: 8, sub O. foetida var. broteri; Carlón & al., 2005:  37 ["arenales costeros ... castaño anaranjado"], sub O. densiflora). Herbarium Specimens: a) "Herbarium M.J.Y. Foley / Flora of Spain / Orobanche sanguinea Presl s.l. / Possibly of hybrid origen xO. foetida Poir. ? / Zahara de los Atunes, Cadiz Province, in / coastal dune meadow probably parasitic on / Medicago sp. (also Lotus sp.). Occupied slight / depressions in two places in dunes - but quite / separate habitat from O. densiflora Salzm. which / is abundant here. Whole plant and stigma lobes / deep wine red / nº. 243 Leg. M.J.Y. Foley / Date 26/27-IV-1993 Det. M.J.Y. Foley / This plant is under further investigation and is / possibly a subsp. nov. - MJYF (1993). // b) Holotype Orobanche crinita  Viv. var. occidentalis M.J.Y. Foley // c) Holotype Orobanche crinita Viv. subsp. occidentalis M.J.Y. Foley // d) Orobanche crinita Viviviani subsp. / occidentalis M.J.Y. Foley / Fide M.J.Y. Foley 4-2-1998 // e) Orobanche sanguinea / of possible hybrid origen xO. foetida Poir. / Determinavit M.J.Y. Foley MJYF I/1995" (E 00053427)

= Orobanche densiflora var. tarifae A. Pujadas in Acta Bot. Malacitana 33: 363 [1 online] (2008 [Dec. 2008 (online, 14 Oct. 2008, vide Vienna Code, Art. 29)]). Syn.: = Orobanche densiflora Salzm. ex Reut. (1847) [red flowering forms with yellow stigmas]. Ind. loc.: "Holótipo: Cádiz [Andalucía / Andalusia, España / Spain], Tarifa, Playa del Cañuelo, c. Faro Camarinal, 36º 5,383 N 5º 48,326 W, 9 m, 10-V-2008, A. Pujadas, COA45361". Obs. "excepto el estigma que es amarillo" (cf. A. Pujadas, 2008: 1; Carlón & al., 2005: 37 "estigmas amarillos, rojos... o de un castaño anaranjado"; Carlón & al., 2005: 37, 61 f. 4, 64 "se ha determinado en las dos formas la mismísima secuencia ITS —cf. Figura 4. [The ITS sequence for yellow and red flowering forms has been studied by Schneeweiss, the results indicate that it is the same for both sympatric forms (61 f. 4, 64)]"). SALA 136763 Isotype

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? = Orobanche densiflora var. genuina f. macraixma (J.A. Guim.) Beck in Engl., Pflanzenr. 96: 195 (1930) ["Forma 3. macraichma"]. Syn.: Orobanche densiflora var. macraixma Guim. in Brotéria 3: 144[-148] (1904)

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Misapplied names:

Orobanche crinita auct. pl., non Orobanche crinita Viv., Fl. Cors. Prodr.: 11 (1824)

Orobanche crinita sensu Rchb., Iconogr. Bot. Pl. Crit. 7: 43, t. 687 f. 922 (1829), non Orobanche crinita Viv., Fl. Cors. Prodr.: 11 (1824). Syn.: = Orobanche densiflora Salzm. ex Reut. (1847) [sec. Reichenbach, 1829: 43, "O. densiflora Salzm. pl. hisp. tingit. exsicc."; Reuter, 1847: 19; Cosson, 1849-1852: 42; Nyman, 1854-1855: 130; Willkomm, 1870: 622; Guimarães, 1904: 143; Beck, 1930: 193; (!)]. Ind. loc.: "Hist. 922. in sabulosis maritimis circa Tanger [Tangier, Région de Tanger-Tétouan, Morocco / Marruecos]; Salzmann."

? – Orobanche densiflora sensu Bertol., Fl. Ital. 6: 444-445 (1846), non Orobanche densiflora Salzm. ex Bertol., Fl. Ital. 6: 444 (1846). Ind. loc.: "Ann. Habui ex Castel Sardo [Castelsardo / Castel Sardo / Caltheddu / Casteddu Sardu, Province of Sassari, Sardegna / Sardinia / Cerdeña, Italia / Italy] in Sardinia septentrionali ab Eq. Prof. Vivianio, qui confundebat eam cum sequente". Obs. "Corolla parva, tubulosa ... superne recurva, pallide flavescens ... Stigmatis  ... purpurei" (cf. Bertoloni, 1846: 441); "Orobanche densiflora Salzm. ex Reut. (in Candolle, Prodr. 11: 19. 1847) was reported from Castelsardo (Bertoloni 1846), based on a specimen collected by Viviani (BOLO!) that belongs to O. litorea Guss. Moris (1858-59) had seen it in Viviani’s own herbarium, mixed with O. crinita. Beck (1890) considered "O. densiflora" in the sense of Moris and Bertoloni as a variant of O. minor Sm. Nevertheless, O. densiflora is, although doubtfully, recorded for Sardinia by Chater & Webb (1972), and fully so by Conti & al. (2005)" (cf. Domina, & Arrigoni, 2007: 134)

Orobanche densiflora sensu Willk. in Willk. & Lange. Prodr. Fl. Hispan. 2(3): 622 (1870) [p.p.], non Orobanche densiflora Salzm. ex Willk. in Willk. & Lange. Prodr. Fl. Hispan. 2(3): 622 (1870). Obs. vide Orobanche foetida Poir.

Orobanche foetida auct. [saltem p.p.], non Orobanche foetida Poir., Voy. Barbarie 2: 195 (1789) (cf., e.g., Willkomm, 1870: 622)

Orobanche foetida var. broteri sensu Pujadas & al. (2003: 740, 742), p.p., non Orobanche foetida var. broteri (J.A. Guim.) Merino, Contr. Fl. Galicia 2: 49 (1897). Syn.: = Orobanche crinita var. occidentalis M.J.Y. Foley (2001); – Orobanche densiflora Salzm. ex Reut. (1847) [red flowering form with orangish-brown stigma]. Obs. Carlón & al. in Documentos Jard. Bot. Atlántico (Gijón) 3: 37 (2005)

Orobanche foetida var. foetida sensu Pujadas & al. (2003: 740, 742), p.p., non Orobanche foetida Poir., Voy. Barbarie 2: 195 (1789). Syn.: Orobanche densiflora Salzm. ex Reut. (1847) [red flowering form with orangish-brown stigma]. Obs. Carlón & al. in Documentos Jard. Bot. Atlántico (Gijón) 3: 37 (2005)

Orobanche foetida var. lusitanica sensu Pujadas & al. (2002: 434), p.p., non Orobanche foetida subsp. lusitanica [Brot. ex] Cout., Fl. Portugal: 568 (1913). Syn.: = Orobanche foetida var. broteri sensu Pujadas & al. (2003: 740, 742), p.p., non (J.A. Guim.) Merino (1897); Orobanche densiflora Salzm. ex Reut. (1847) [red flowering form with orangish-brown stigma]. Obs. Carlón & al. in Documentos Jard. Bot. Atlántico (Gijón) 3: 37 (2005)

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Ind. loc.: "in sabulosis maritimis circa Tanger [طنجة / Tánger / Tanja / Tangier, Région de Tanger-Tétouan, Morocco / Marruecos]; Salzmann." (Reichenbach, 1829, sub O. crinita sensu Rchb., Iconogr. Bot. Pl. Crit. 7: 43, t. 687 f. 922); "In arenosis maritimis regni Maroccani prope Tingidem [طنجة / Tánger / Tanja / Tangier] (Salzm.! Schousb. in h. Pavon! nunc Boiss.)" (Reuter, 1847: 19; Foley, 2001: 54).

Typus / Type: W 1889-0082394 [Herb. Mus. Palat. Vindob. / Collectio Reichenbach fil.] (Reuter, 1847: 19, "Salzm.! pl. tingit. exsicc."). Lectotypus / Lectotype: "Orobanche densiflora S. Cat. / M. Salzmann 1839 / Tanger in sabulosis maritimis" (G) (Domina & Soldano, 2009). Isotypus: a) "Orobanche densiflora Slzm. cat. / Tanger, in sabulosis maritimis", b) "Herb. Mus. Paris / Orobanche densiflora / Salzm.! / Tanger  M. Salzmann" (P 02978276; rev. Ó. Sánchez Pedraja Nov 2011). Typus: a) "Orobanche: / spicâ conicâ floribus minoribus / ... / Tanger / Herb. Pavon", b) "Orobanche densiflora Salzm.! / Reuter in DC prod. / Orobanche crinita Rchb. pl. crit. ..." (G 00383153 - hb. Boissier; rev. Ó. Sánchez Pedraja May 2016).

Designated by: M.J.Y. Foley, 1995-02 [in sched.]. Domina & Soldano in 104° Congr. Soc. Bot. Italiana (2009).

Host: usually on Lotus creticus L. (Leguminosae), more rarely on other Leguminosae species (e.g., Ononis variegata L.) (Annotated Checklist of Host Plants of Orobanchaceae, accessed Aug 2009).

Comments: cariological and genetic studies suggest that Orobanche densiflora were not connected with Orobanche minor group. (Román & al., 2003; Schneeweiss & al., 2004; Carlón & al., 2005: 62), contrary to what some authors seem to think (e.g., Beck, 1930; Uhlich & al., 1995: 11). Moreover, O. densiflora is sometimes confused with O. foetida, it can be differentiated by an slightly aromatic odor, which is smoother and more pleasant than in the case of O. foetida (Carlón & al., 2005: 40).

Carlón & al. in Documentos Jard. Bot. Atlántico (Gijón) 3: 36, 37, 38 t. 13, 39 t. 14, 40, 61 f. 4 ["Phylogenetic relationships"], 64 (2005); Documentos Jard. Bot. Atlántico (Gijón) 6: 13, f. 2 [phylogenetic relationships] (2008).

Chromosomatic number: n = 38 (Palomeque & Sañudo, 1981 [Cádiz, Spain]); 2n = 76 (Delay & Petit, 1971; Schneeweiss & al., 2004).

Ic.: a, inflorescences [yellow flowering forms; Tarifa, Spain]; b, bract, calyx-segments and pistil (ib.); c, inflorescences [yellow and red flowering (red stigma) forms; ib.]; d, red flowering form with red stigma (ib.); e, opened corolla showing stamens (ib.); f, corolla, in partial long-section showing pistil (ib.); g, bract and calyx-segments (ib.); h, shoot [yellow flowering form, ib.]; i, inflorescences [yellow and red (yellow stigma) flowering forms; Tarifa-Caraminal, Spain]; j, inflorescences [red flowering forms with yellow stigmas; ib.]; k, flowers [yellow and red flowering forms with yellow and red stigmas; Tarifa, Spain]; l, P 03428429 - herb. E. Cosson, 18, Tánger [loc. class.]; m, Reichenbach, Iconogr. Bot. Pl. Crit. 7: 43, t. 687 f. 922. 1829 (Tangier, Morocco, sub O. crinita); n, Beck in Biblioth. Bot. 19: t. 4 f. 71 (1890); o, Orobanche foetida (10504) vs O. densiflora (13488) vs O. crinita (11204); p, Orobanche foetida (13482) vs O. densiflora (11759) [bracts and calyces].

Distribution: Mediterranean Region distribution; Iberian Peninsula distribution.

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Specimens examined and literature records.

Algeria / الجزائر‎ / Argelia / Algérie [unconfirmed, although it was cited by Battandier (1890), this mention seems to be erroneous]

Oran: "Oran (Munby)" (Battandier, 1890: 660) [unconfirmed (Beck 1930: 300, sub O. sanguinea f. loriformis)].

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Italy / Italia / Italien [unconfirmed]

Sardinia / Sardegna / Cerdeña

Sardinia: "Ann. Habui ex Castel Sardo [Province of Sassari] in Sardinia septentrionali ab Eq. Prof. Vivianio, qui confundebat eam cum sequente" (cf. Bertoloni 1846: 440, sub O. densiflora); "Orobanche densiflora Salzm. ex Reut. (in Candolle, Prodr. 11: 19. 1847) was reported from Castelsardo (Bertoloni 1846), based on a specimen collected by Viviani (BOLO!) that belongs to O. litorea Guss. Moris (1858-59) had seen it in Viviani’s own herbarium, mixed with O. crinita. Beck (1890) considered "O. densiflora" in the sense of Moris and Bertoloni as a variant of O. minor Sm. Nevertheless, O. densiflora is, although doubtfully, recorded for Sardinia by Chater & Webb (1972), and fully so by Conti & al. (2005)." (Domina, & Arrigoni, 2007: 134).

Sicily / Sicilia / Sizilien

Sicilia: the Sicilian records may be due to confusion between O. densiflora and littoral specimens with dense inflorescences of O. sanguinea C. Presl [= O. crinita Viv. = O. sanguinea var. maritima Lojac.]. E.g., "FI / Plantae siculae rariores / 463. Orobanche sanguinea Presl / del. Prag. p. / O. densiflora Salzm. (Auch) / var. maritima mihi monogr. / orob. ined. / In sylvaticis herbosis maritimi / mayo 1881 ... in Gallo parte di / Sferracavallo / leg. M. Lojacono." (Domina & Mazzola, 2007: 226-227, sub O. sanguinea f. crinita (Viv.) Beck).

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Malta [unconfirmed]

Malta: "a) O. densiflora Salzm. / f. melitensis m. / differt a typo dentibus calycis / partium alte connatis, corolla subcopiose / glandulosa, glabrescens. / Ein kleines Exemplar / behalten! / Sehr interessanter Standort!" // b) Stéphen Sommier – Plantae Melitenses / Orobanche densiflora Salzm. / f. melitensis m. / Beck / Insula Melita (hodie Malta) / Marfa – in litore arenoso. / 4 Maii 1907 legi / ex albo-flavescens / 55 // c) Holotypus" (FI 002514); "Auf der Insel Malta an der Küste bei Marfa, im Mai (Sommier in der Var. β!)" (cf. Beck, 1930: 194); Malta (cf. S. Mifsud, sub O. densiflora f. melitensis Beck; accessed, 2009).

N. B. "Lo studio del materiale originale, conservato a Firenze (FI!), ha permesso di riferire questo taxon a O. sanguinea C. Presl. ... escludono la presenza di Orobanche densiflora Bertol. dal territorio italiano e da Malta." (cf. Domina & Soldano, 2009).

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Morocco / المغرب‎, / Marruecos / Marokko

Grand Casablanca [unconfirmed]

Casablanca: "Casa Blanca [Casablanca / Ad Dār al Baydā', N 33° 35' 24'' W 7° 36' 36''] !" (cf. Ball, 1877-1878: 604; Beck, 1890: 233).

Marrakech-Tensift-Al Haouz [unconfirmed]

Iles Purpuraires: "B. Balansa, Plante du Maroc, 1867 / Orobanche foetida Desf. (Coss.) / Stigmates d'un jaune rougeâtre. / Parasite sur Lotus Salzmanni [Lotus creticus L.] / Mogador  Mai" (P 03428652 - hb. E. Cosson; rev. Ó. Sánchez Pedraja Oct 2014 [need to be confirmed, see O. foetida]); idem (P 02978306 - hb. E. Drake; rev. Ó. Sánchez Pedraja Oct 2014 [need to be confirmed, see O. foetida]); ibidem (P 02978374; rev. Ó. Sánchez Pedraja Oct 2014 [need to be confirmed, see O. foetida]); ibidem (P 03428420; rev. Ó. Sánchez Pedraja Oct 2014 [need to be confirmed, see O. foetida]).

Tangier-Tetouan

Tanger-Assilah: "Near Tangier. M. Durand. 1803. N.º 32 // Det.-Rev. M.J.Y. Foley 2.iii.2000" (LINN-HS1087-2, O. densiflora in Jstor; accessed Nov 2014); "Orobanche densiflora mihi [Salzmann] / in sabulosis maritimis c. Tanger // O. crinita Viv." (W 1889-0082394; type of O. densiflora Salzm., det. M.J.Y. Foley II-1995); "Orobanche densiflora Salzm. / Tanger in sabulosis maritimis / ..." (MZM 5963, MZM5963a); "Orobanche densiflora / Tanger in sabulosis maritimis" (MPU 021201, Isosyntype of O. densiflora in Jstor; accessed, Nov. 2014); "Orobanche densiflora Salzmann / Salzmann 1835. / Tanger in sabulosis maritimis // Orobanche densiflora S. / Tanger in sabulosis maritimis" (MPU 021202, Isosyntype of O. densiflora in Jstor; accessed, Nov. 2014); "Orobanche densiflora mihi / Herb. Salzmann / in sabulosis maritimis c. Tanger. Majo" (MPU 021203, Isosyntype of O. densiflora in Jstor; accessed Nov 2014); "Herbarium Regium Monacense / Orobanche densiflora / ... / Tanger in sabulosis maritimis / Legt. et ... Salzmann / Herb. Zuccarinii" (M 0185486, Isosyntype of O. densiflora in Jstor; accessed Nov 2014); "Orobanche foetida / in sabulosis maritimis pr. Tanger. // Orobanche foetida Desf. Salzmann! / Tanger  M. Salzmann." (P 02978383; rev. Ó. Sánchez Pedraja Nov 2014); a) "Orobanche: / spicâ conicâ floribus minoribus / ... / Tanger / Herb. Pavon", b) "Orobanche densiflora Salzm.! / Reuter in DC prod. / Orobanche crinita Rchb. pl. crit. ..." (G 00383153 - hb. Boissier; rev. Ó. Sánchez Pedraja May 2016, type of O. densiflora); "In arenosis maritimis regni Maroccani prope Tingidem [loc. class., Tangier, N 35° 46' 48'' W 5° 48' 36''] (Salzm.! Schousb. [see O. foetida] in h. Pavon! nunc Boiss.)" (cf. Reuter, 1847: 19, sub O. densiflora; Beck, 1890: 233); "in sabulosis maritimis circa Tanger; Salzmann." (cf. Reichenbach, 1829, sub Orobanche crinita sensu Rchb., Iconogr. Bot. Pl. Crit. 7: 43, t. 687 f. 922); "Tanger (Salzm. Exsicc.)!   Ad ostium fluminis Tetuanensis (1851)!   Specimen a G. Maw fere eodem loco lectum (1869) in Herb. Kew.!" (cf. Ball, 1877-1878: 604; Beck, 1890: 233; K - Hookerianum 0004251200 [!, this specimen was probably collected by Salzmann in Tanger (Morocco)]); "Iter algeriensi-hispanicum. / Orobanche densiflora Desf. / Mauritania / Tanger / Anno 1849.  Leg. Boissier & Reuter." (W - Rchb. 1889-0082393; Det. M.J.Y. Foley, 2.1995, ut O. densiflora); "Cabo Espartel, Tánger" (cf. Pau., 1924: 371, [unconfirmed]); a) "Herb. Mus. Paris / Orobanche densiflora / Salzmann! / Tanger / M. Salzmann", b) "Orobanche densiflora Slzm. cat. / Tanger, in sabulosis maritimis" (P 02978276; rev. Ó. Sánchez Pedraja Nov 2011, isotypus); "Nº 226 / Orobanche foetida Desf.! Boiss.! / non DC!  / ... / Tanger sables maritimes. Sur le / Lotus arenarius [Lotus creticus L.?]!  mayo 1849 / Reuter" (P 03428648 - hb. E. Cosson; rev. Ó. Sánchez Pedraja Oct 2014 [only one specimen close to its label]); "Orobanche foetida Desf.! / Tanger arenis maritimes, ad rad. / Lotus arenarii [L. creticus L.?] mayo 1849 / Reuter" (P 03428647 - hb. E. Cosson; rev. Ó. Sánchez Pedraja Oct 2014 [the sheet contains two specimens, one of these appears to be O. densiflora but the other (the right of the pair) is from J. Ball's 1851 and this seems to correspond to O. foetida]); a) "Orobanche densiflora Salzm. / Tanger, in dunis arenosis / in radice Loti Salzmnnii [L. creticus L.] / mayo 1849 / Reuter", b) "Salzmann. Plantae Tingitanae exsiccatae. Distr. 1824 / Orobanche densiflora Salzm. / Tanger, in sabulosis maritimis / 1835", c) "Boissier et Reuter / Iter Algeriensi-Hispanicum. / Orobanche densiflora Salzm! / O. crinita Rchb. non Viv. / Tanger / sur les dunes de sables blanc / pres la mer. parasite sur un / Lotus (L. Salzmanni Boissier et Reut. [L. creticus L.]) / leger. Boissier et Reuter / Maio 1849." (P 03428429 - Herb. E. Cosson, 18; rev. Ó. Sánchez Pedraja Nov 2011 [the sheet contains three specimens (each of them next to its label) and all of them correspond to O. densiflora]); "Orobanche foetida Desf.! non DC. / Tanger arenis maritimes / sur le Lotus arenarius [L. creticus L.?]! april 1849 / Reuter" (P 02978385; rev. Ó. Sánchez Pedraja Nov 2011).

  [N.B. O. densiflora, O. foetida, O. sanguinea [incl. O. crinita] and O. variegata are four relatively close species which are probably all present on the northern coast of Morocco and Algeria (cf., e.g., Ball, 1878: 604; Pau, 1924: 371; K 000425199-202 [mixed gathering of different specimens from Algeria and Morocco]). This could be a possible source of misidentifications, specially with the less typical specimens].

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Portugal [unconfirmed]

Algarve (Ag)

Faro District: Castro Marim [N 37° 13' 0'' W 7° 26' 0''] (cf. Gandoger, 1901: 409; Gandoger, 1917: 273; Beck, 1930: 194) [unconfirmed]; "Aljezur ...14-4-1994" [N 37° 19' 0'' W 8° 48' 0'', yellow flowering form! [parasitic on Ononis natrix (cf.)] (Kreutz, 1995: 13).

N. B. Thorogood has only seen robust yellow forms of O. foetida (pers. comm., 2010).

Baixo Alentejo (BAL)

Beja District: "N.º 20 Herbarium lusitanicum / J. Daveau / Orobanche densiflora Salzm / Jachères près de Serpa [Beja District] / Mai 1882" (P 04419811, rev. Ó. Sánchez Pedraja Nov 2011 [the sheet contains four specimens (each of them next to its label), three of them correspond to Orobanche densiflora from Cádiz (Spain) and the fourth (the Daveau's specimen) seems to correspond to O. gracilis from Serpa (see also O. rapum-genistae)]).

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Spain / España / Espagne / Spanien

Andalusia / Andalucía

Cádiz (Ca): "Civis Europae maxime australis et Barbariae" (Reichenbach, 1829: 43, sub Orobanche crinita); a) "E. Bourgeau, Pl. d'Espagne, 1849. / 376. Orobanche densiflora, Salzm.! / (Reuter.) / Dans les sables mouvants entre Santa Cathalina et Puerto / Santa Maria, parasite sur le Lotus Creticus. / 17 Mars.", b) "Herb. Mus. Paris / O. densiflora Salzm. / Baetica" (P 04419805; rev. Ó. Sánchez Pedraja, Nov. 2011); ibidem (P 04419809; rev. M.J.Y. Foley, 1998; Ó. Sánchez Pedraja, Nov. 2011); (P 04419806 - herbier E. Drake; rev. Sánchez Pedraja, Nov. 2011); ibidem (P 04385361; rev. Ó. Sánchez Pedraja, Nov. 2011 [only a small specimen attached to the host-plant, the bigger specimen is O. gracilis]); ibidem (P 04410456 - Herbier L. Giraudias ex Ecole Normale Sup.; rev. Ó. Sánchez Pedraja, Nov. 2011); "Orobanche densiflora Salz. / Andalusie, Bourgeau" (P 04419804; rev. Ó. Sánchez Pedraja, Nov. 2011 [the sheet contains two specimens]); "In arenosis maritimis agri Gaditani prope Puerto Santa Maria, in radicibus Loti Cretici parasítica! (E. Bourgeau, pl. Esp. 1849, n. 576). Cette espèce, nouvelle pour l'Europe, a été déterminée par M. Reuter, qui a bien voulu se charger de l'étude des plantes de la famille des Orobanchées recueillies par M. Bourgeau." (cf. Cosson, 1849: 42, sub O. densiflora) ; "Cadix" (cf. Nyman, 1854-1855: 130; Nyman, 1881: 559, "Exs. Bourg. hisp. a. 1849. 376"); "Baet. (pr. Pto. de Sta. Maria [N 36° 35' 38'' W 6° 13' 58''], Bourg.)" (cf. Willkomm, 1870: 622; Colmeiro, 1888: 276; Beck, 1890: 233; Gandoger, 1917: 273); a) "Reise durch Spanien und Portugal / Orobanche densiflora Salz. / inter Palmones et Algeciras /  17/4 1876 M. Winkler", b) "Reise durch das südliche Spanien 1873 / Orobanche densiflora Salzm. / Palmones, p Algeciras / in radice Leguminosam / 10 May   M. Winkler" (P 04419811; rev. Ó. Sánchez Pedraja, Nov. 2011 [the sheet contains four specimens (each of them next to its label), three of them correspond to Orobanche densiflora from Cádiz and the fourth (the Daveau's specimen) seems to correspond to O. gracilis from Serpa (see also O. rapum-genistae)]); "entre Algésiras et San-Roque (Winkl.), à Algésiras (Rev.)" (cf. Debeaux & Dautez, 1889: 154, sub densiflora); "Reise durch das südliche Spanien 1873. / Orobanche densiflora Salzm. / Algeciras, Palmones, in Loto / cretico [Lotus creticus L.] et Medicago marina [Medicago marina L.] / 10.5.   R. Fritze." (COI 00056875, rev. Ó. Sánchez Pedraja, March 2012); "pr. Palmones [N 36° 11' 0'' W 5° 26' 0''] et Algeciras [N 36° 7' 39'' W 5° 27' 14''] ad radic. Loti cretici [Lotus creticus L.] et Medicag. marinae [Medicago marina L.] (Fritze!, Winkler! 1873, Reverch., 1887" (cf. Willkomm, 1893: 187; Beck, 1890: 233; Beck, 1930: 194); "Sanlúcar de Barrameda [N 36° 47' 0'' W 6° 21' 0''] (Colm.)" (cf. Colmeiro, 1888: 276); "Algeciras, Palmones in Loto cretico et Medicago marina, 10-V-1873, R. Fritze, MAF 36183 & COI-Hº Willkomm (citado en Willkomm, 1893: 187)" (cf. Pujadas & al., 2007: 20); In itinere hispanico-lusitanico / Hispania: In radicibus Loti cretici pr. Algeciras, E. Hackel, s.n. [1876-04-17] (W 0021305); Cerca de Sanlúcar [de Barrameda], [29SQA37] ... Puerto de Santa María, 29SQA45 ... entre Algeciras [N 36° 7' 20'' W 5° 26' 40''] y San Roque, 30STF70 (cf. Pérez Lara, 1889 in Anthos, accessed, 8 Mar. 2010); Orobanche densiflora, "between La Paloma and Bolonia near Tarifa", 1975/04/22 (cf. Stridvall L. & A., Plant Galleries, accessed, 8 Mar. 2010); "Litoral gaditano, costa de Algeciras" (cf. Valdés, 1987: 554); "Zahara de los Atunes, 1996, Foley, s.n. (v.v.); Punta Paloma, 1973, Smythies 793 (LTR!) ... forma de coloración rojiza" (cf. Foley, 2001b: 54-55); "O. densiflora Salzm. ex Reut. (yellow-flowered form) - AY209243: Spain, Andalucía, province of Cádiz, coast near Conil de la Frontera, 0-5 m, on Lotus creticus (Fabaceae), GS & HW (WU: S&T 8733); AY960725: Spain, Andalucía, province of Cádiz, coastal area of the Paloma Baja E to NE of Punta Paloma, 0-40 m, on Lotus creticus (Fabaceae), WG (Herb. Gutermann 37440); O. densiflora Salzm. ex Reut. (red-flowered form) - AY960727: Spain, Andalucía, province of Cádiz, coastal area of the Paloma Baja E to NE of Punta Paloma, 0-40 m, on Lotus creticus (Fabaceae), W. Gutermann (Herb. Gutermann 37439)" (cf. Carlón & al., 2005: 64); Tarifa, dunas de Bolonia, 30STE4997, 5 m, dunas, G. Gómez Casares & G. Moreno Moral MM0011/2004, 15/02/2004 (herb. Sánchez Pedraja 11698 [yellow and red flowering forms]); Tarifa, dunas de Punta Paloma, 30STE5694, 15 m, dunas, G. Gómez Casares & G. Moreno Moral, 15/02/2004 (obs.); Cádiz, Junto al Ventorrillo del Chato —playa de Cortadura—, 29SQA4541, 2 m, sobre (!) Lotus creticus L. (Leguminosae), dunas, G. Gómez Casares & G. Moreno Moral MM0036/2004, 09/04/2004 (herb. Sánchez Pedraja 11744 [yellow flowering forms]); Tarifa, dunas de Zahara de los Atunes, 30STF4402, 2 m, sobre Lotus creticus L. y Medicago marina L. (Leguminosae), dunas, G. Gómez Casares & G. Moreno Moral MM0040/2004, 09/04/2004 (herb. Sánchez Pedraja 11751); Tarifa, dunas de Punta Paloma, 30STF5694, 20 m, sobre (!) Ononis variegata L. (Leguminosae), dunas, G. Gómez Casares & G. Moreno Moral MM0046/2004, 10/04/2004 (herb. Sánchez Pedraja 11758); Tarifa, dunas de Punta Paloma, 30STF5694, 20 m, sobre (!) Lotus creticus L. (Leguminosae), dunas, G. Gómez Casares & G. Moreno Moral MM0047/2004, 10/04/2004 (herb. Sánchez Pedraja 11759 [yellow and red flowering forms]) (cf. Carlón & al., 2005: 39); Tarifa, dunas de Punta Paloma, 30STF5694, 20 m, sobre (!) Lotus creticus L. (Leguminosae), dunas, G. Gómez Casares & G. Moreno Moral MM0048/2004, 10/04/2004 (herb. Sánchez Pedraja 11760 [red flowering forms]); Tarifa, dunas de Punta Paloma, 30STF5694, 20 m, sobre (!) Lotus creticus L. (Leguminosae), dunas, G. Gómez Casares & G. Moreno Moral MM0049/2004, 10/04/2004 (herb. Sánchez Pedraja 11761 [yellow flowering forms]); Tarifa, bajo el faro Camarinal [pr. "Urb. Atlanterra"] —Zahara de los Atunes—, 30TSF4797, 20 m, sobre Lotus creticus L. (Leguminosae), dunas, G. Gómez Casares & G. Moreno Moral MM0091/2004, 22/05/2004 (herb. Sánchez Pedraja 11846 [yellow and red flowering forms]); Tarifa, Zahara de los Atunes, 30TSF4401, 2 m, sobre Lotus creticus L. (Leguminosae), dunas, G. Gómez Casares & G. Moreno Moral, 22/05/2004 (obs. [yellow flowering forms]); Tarifa, bajo el faro Camarinal [pr. "Urb. Atlanterra"] —Zahara de los Atunes—, 30TSF4797, 20 m, sobre (!) Lotus creticus L. (Leguminosae), dunas, G. Gómez Casares & G. Moreno Moral MM0092/2004, 22/05/2004 (herb. Sánchez Pedraja 11847) (cf. Carlón & al., 2005: 38); Spanien, prov. Càdiz, Atlantik-Küste NW von Tarifa, SE von Zahara de los Atunes, N- Ende des Strandes SE von Cabo de Plata; 5-10 m S. M.; 36°06'04" N / 05°49'05" W, Vitek, E. 06-0014, 2006-04-13 (W 2006-0017067); "I think Orobanche densiflora, maybe ... Cádiz ... (mapa) ... 19 de abril de 2006" (cf. Harry, R. in www.flickr.com, accessed 3 Mar. 2010); Barbate, en el Parque Natural de la Breña y Marismas de Barbate, 30STF3408, 90 m, junto a Lotus cytisoides [Lotus creticus L.], suelos arenosos, J.A. García Rojas, G. Gómez Casares & G. Moreno Moral, 26/03/2007 (obs.); "Orobanche densiflora, fotografiado en el tómbolo de Trafalagar en Barbate", Junio 5, 2006 (cf. Juanjo in Hablando en verde, accessed, 8 Mar. 2010); "Puntualmente abundante en cordones dunares y arenas interiores de origen marítimo. Principalmente parasita a Lotus creticus aunque en esta ocasión se instaló sobre Ononis dentata. La foto se tomó en la primavera de 2007 en Cabo Roche, Conil de la Frontera (Cádiz)" (R. Sánchez Carrión in Fotonatura, www.fotonatura.org, accessed, Mar. 2010); O. densiflora, "Arenales costeros / ... / Costa del Estrecho, CA [Cádiz]" (cf. Quiles Hoyo; accessed, 23 May 2009); "pinares de Barbate dentro de P.N.La Breña, el dia 10 de Mayo de 2008 / ... / Orobanche densiflora. Jopo. Planta parásita de leguminosas. Abundante en cordones dunares y arenas interiores de origen marítimo. Principalmente parasita a Lotus creticus" (cf. Joaquín in Tercera Ruta Botánica del Grupo Senderista el Batolito, Picasa, accessed, Mar. 2010); municipality of Barbate, W of the Yerbabuena beach, near Barbate village, 30STF3408, 20 m, next to Lotus creticus L. and Ononis natrix subsp. ramosissima, G. Moreno Moral, 2 Apr. 2010 (obs.); ib., Pinar de Barbate, near Barbate village, 30STF3409, 95 m, slope on the road, Moreno Moral, 2 Apr. 2010 (obs.); municipality of Tarifa, Punta Paloma dunes (at the Macotilla place), near Casas de las Palomas, 30STE5694, 20 m, next to Lotus creticus L., Moreno Moral MM0024/2010, 2 Apr. 2010 (herb. Sánchez Pedraja 13487) [we only gathered a red flowering specimen, but we could see hundreds of specimens of this species in these dunes]; municipality of Barbate, the Calderas beach, near Zahora village, 29SQA6510, 5 m, parasitic on (!) Lotus creticus L. in tertiary dunes, G. Moreno Moral MM0025/2010, 3 Apr. 2010 (herb. Sánchez Pedraja 13488) [very abundant in some places of the Calderas beach. In this beach I have seen dry specimens of the last year and I have also seen a great number of flowering and sprouting specimens of this year. Here, the stream of San Ambrosio will sink into the sand just before emptying into the sea. I went to this place to investigate the following record of Orobanche var. foetida: "estuario arroyo San Ambrosio, Caños de Meca, 20-II-2002, E. Sánchez Gullón & P. Weickert, COA 37523 (cf. Pujadas & al., 2007: 108)". I found no specimen of this species, but the estuary was not surveyed extensively (see O. foetida)]; "Cádiz, Tarifa, Playa del Cañuelo, c. Faro Camarinal, 36º 5,383 N 5º 48,326 W, 9 m, 10-V-2008, A. Pujadas, COA45361" (cf. A. Pujadas, 2008: 1, holótipo of O. densiflora var. tarifae, SALA 136763 Isotype in Jstor; accessed Nov 2014).

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Tunisia / تونس / Tunisie / Túnez (Raffaelli & Ricceri, 1989; Le Floc’h & al., 2010)

Nabeul / Nābil: "Ce taxon ibéro-maghrébin, à fleurs jaune clair, a été relevé, pour la première fois en Tunisie entre El Haouaria et Ras Adar (Cap Bon), le 13.04.1986, par Raffaelli & Ricceri [Candollea 41: 99 (1989)]" (Le Floc’h & al., 2010: 273).

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