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Accepted Name - Basionymum / Basionym - Synonyms - Ind. loc. [Indicatio locotypica (type locality)] - Lectotypus [Lectotype] - Designated by - Host - Comments - Chromosomatic number - Ic. [Images] - Distribution - Specimens Examined - References - Authors - Cite us ISSN: 2386-9666 - Last Updated: 10 August 2023 |
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Accepted Name: Orobanche hederae [Vaucher ex] Duby, Bot. Gall.: 350 (1828 [12-14 Apr 1828]) ["10. O. Hederae (Vauch. orob. p. 56. t. 8.)"] Synonyms: Homotypic synonyms: ≡ Orobanche du Lierre Vaucher, Monogr. Orobanch.: 56, t. 8 (1827), nom. inval. [not validly published] (Reichenbach f. 1862: 103, sub O. Hederae Duby). Syn: ≡ Orobanche hederae [Vaucher ex] Duby, Bot. Gall.: 350 (1828) (Reuter 1847: 28, sub O. Hederae Duby); ≡ Orobanche hederae-helicis [Vaucher ex] F.W. Schultz in Bot. Lit.-Blätt. 5: 500 (1830); ≡ Orobanche hederae [Vaucher ex] Des Moul. in Ann. Sci. Nat., Bot., ser. 2, 3: 80 (1835); ≡ Orobanche vaucherii Noulet, Fl. Bass. Sous-pyrén.: 487 (1837); ≡ Orobanche hederae [Vaucher ex] Grenier in Grenier & Godron, Fl. France 2(2): 640 (1853); ≡ Catodiacrum luteum Dulac, Fl. Hautes-Pyrénées: 370 (1867); ≡ Orobanche hederae [Vaucher ex] Dulac, Fl. Hautes-Pyrénées: 370 (1867), pro syn; ≡ Orobanche hederihaerens St.-Lag., Cat. Fl. Bassin Rhône: 611 (1883); ≡ Orobanche hederifixa St.-Lag. in Cariot, Etud. Fl. ed. 8(2): 651 (1889). Ind. loc.: "Cette singulière Orobanche se trouve sur les collines qui s'étendent de Gênes à Savone [Genoa / Genova to Savone / Savona, Liguria, Italy]. Je l'ai cueillie en 1825, sur les hauteurs qui dominent cette dernière ville, et dans les maisons de campagne du Ponente. Je l'ai revue l'année suivante en grande abondance à Gênes même, dans les jardins Négri. Elle vient d'être retrouvée par M. Charles Des Moulins, dans le Périgord [Dordogne, Nouvelle-Aquitaine, France] et aux environs de Libourne [Gironde, Nouvelle-Aquitaine, France]. Elle fleurit dans le mois de Juin." (Vaucher 1827: 57) ≡ Orobanche hederae-helicis [Vaucher ex] F.W. Schultz in Bot. Lit.-Blätt. 5: 500 (1830) ["18. O. Hederae Helicis Tab. VIII."], nom. superfl. (Beck 1930: 219) ≡ Orobanche hederae [Vaucher ex] Des Moul. in Ann. Sci. Nat., Bot., ser. 2, 3: 80 (1835) ["5. Orobanche Hederae Vauch. ... O. Hederae Vauch. Monogr. Orob. p. 56, n° 18, tab. 8.— Duby, Bot. Gall. 1, p. 350, n° 10."], nom. superfl. Ind. loc.: "à Lanquais comme aux environs de Libourne et de Bordeaux [France]" (Des Moulins 1835: 80) ≡ Orobanche vaucherii Noulet, Fl. Bass. Sous-pyrén.: 487 (1837) ["6. Orobanche de Vaucher. ... O. Vaucherii. Nob."], nom. superfl. (Beck 1930: 219). Ind. loc. "H. Sur les racines du lierre. R. R. Près de Toulouse au Vernet, dans le parc de l'ancien château. C. C. C." ≡ Orobanche hederae [Vaucher ex] Grenier in Grenier & Godron, Fl. France 2(2): 640 (1853) ["O. Hederae (O. du Lierre) Vauch. mon. 56, t. 8"], nom. superfl. ≡ Catodiacrum luteum Dulac, Fl. Hautes-Pyrénées: 370 (1867), nom. illeg. Syn: ≡ Orobanche hederae Vaucher ex Dulac, Fl. Hautes-Pyrénées: 370 (1867) (Beck 1930: 219) ≡ Orobanche hederae [Vaucher ex] Dulac, Fl. Hautes-Pyrénées: 370 (1867), pro syn. ≡ Orobanche hederihaerens St.-Lag., Cat. Fl. Bass. Rhône: 611 (1883), nom. illeg. (Beck 1930: 219) ≡ Orobanche hederifixa St.-Lag. in Cariot, Etud. Fl. ed. 8(2): 651 (1889), nom. illeg. ≡ Orobanche minor var. hederae ([Vaucher ex] Duby) Hausm., Flora von Tirol 2: 651 (1852) ["β Hederae"] (Beck 1890: 260). Syn: "O. Hederae Vauch."; "O. hederae Duby"; "O. minor β Bertol." (Hausmann 1852: 651) ≡ Orobanche hederae var. typica J.A. Guim. in Brotéria 3: 180 (1904) ["α"], nom. inval. [not validly published] (Turland & al. 2018: Art. 24.3) ≡ Orobanche hederae f. typica Beck in Biblioth. Bot. 19: 260, t. 4 f. 83 (1890), nom. illeg. ["Formas...: 1. typica = Orobanche Hederae Duby"], nom. inval. [not validly published] (Turland & al. 2018: Art. 24.3) Heterotypic synonyms: ?
= Orobanche laurina Bonap. ex Bertol., Fl. Ital. 6: 424 (1846) (Reuter
1847:
719;
Saint-Lager
1883:
611;
Rouy
1909:
190
[= Orobanche
minor Sm.]).
Ind.loc.: "Ann. Habui Româ [Rome,
Lazio,
Italy]
ab Ill. Principe de Canino, et Musiniano,
qui detexit parasiticam supra radices Lauri nobilis L." (Nyman
1854-1855: 132;
Grenier 1853: 639, "Elle s'éloigne de l'O. Hederae ...
stigmate pourpré ... Sur les racines du Laurus nobilis; Montpellier
(Godron)";
Reichenbach
fil. 1862: 104, "stigmate purpurato ... Montpellier
[France]
Godron!", t. 181; Beck
1930:
219).
Herbarium Specimens: 1.
"Orobanche laurina Godr.! an Bert. / Jardin du plantes 4 juillet 1804
/ Sur le lierre [Hedera]! ..." – 2. "Orobanche
= Orobanche yuccae Bertol., Fl. Ital. 6: 423 (1846). Ind.loc.: Italy: "Ann. Habui ex sylva Pisana [Pisa, Tuscany] prope mare, et ex horto botanico Pisano, quibus in locis reperta est parasitica supra radices Yucccae aloifoliae L., a Prof. Petro Savio" =
Orobanche glaberrima Guss. ex Reut.
in DC., Prodr.
11: 719 (1847 [25 Nov 1847])
["O. glaberrima
(Guss.!
ined.)"]
(Beck 1890:
260;
Turland
& al. 2018:
Art. 46.6).
Ind.loc.: Italy:
"Prope Neapolim
[Naples, Campania] ad radices Dianthi plumarii Guss. !
... (v. s. in h. Boiss. a cl. Guss. comm.)" (Reuter
1847: 719).
Herbarium Specimens: Holotype:
1.
"Orobanche glaberrima
... [description] ...".
– 2. "Orobanche glaberrima Guss.!
/ Reut. in DC. prod. / [m. Reuter]".
– 3. "O. glaberrima
[m. Reuter]
/
Scripsit = Orobanche helicis Rota, Prospett. Piant. Pav.: 287 (1853) [https://www.ipni.org/n/662402-1] (Beck 1890: 260 "Orobanche Helicis Rota, Flora della provincia di Bergamo [Bergamo, Italy] p. 101 ... Vidi sepc. origin. in herb. Cesati."; Beck 1930: 219). Ind.loc.: Italy: "Bergamo [Bergamo]" = Orobanche cerina W.D.J. Koch ex Facchini, Fl. Südtirols: 76 (1855), pro syn. [sub O. minor]. Ind.loc.: Italy: "Postea satis frequenter crescentem plantam islam ad Hederae Helicis radices in ditione Bulsanensi [Bolzano / Bozen, Trentino-Alto Adige / Südtirol] inveni, quam cum O. minore conjungo.". Herbarium Specimens: "Orobanche cerina / Koch ined. / Ober Bench nordlichsim der Nähe von Botzen [?] / Dr. Facchini" (https://data.biodiversitydata.nl/naturalis/specimen/L.2817685 - L 2817685.jpg, rev. Ó. Sánchez Pedraja Nov 2020) = Orobanche lamyana De Cessac ex Malinvaud, Cat. Esp. Rar. Crit. Env. Limoges: 7 (1859) (Rouy 1909: 185, sub O. amethystea, "plus rarement sur l'Hedera helix (= O. Lamyana de Cessac ap. Malinvaud ... plante plus pâle)") [Orobanche lamyana De Cassac ex Nyman, Consp. Fl. Eur. 3: 562 (1881 [prob. Jul 1881]), view IPNI entry] = Orobanche chaboissaei Lacroix in Bull. Soc. Bot. France 6: 565 (1860) (Beck 1930: 219; https://www.ipni.org/n/662229-1). Ind.loc.: France: "M. Chaboisseau a récolté, sur les racines de l'Angelique des jardins cultivée au séminaire de Montmorillon [Vienne department, in the Nouvelle-Aquitaine region], une Orobanche que M. Schultz rapporte avec hésitation à une espece de la Grèce qu'il a publiée dans le Prodromus" = Orobanche aralioctona Meisn. in Bot. Zeitung (Berlin) 24: 17 (1866) (Beck 1930: 219). Ind. loc. "Im Juli dieses Jahres (1865) während ich längere Zeit abwesend war, bemerkte der Obergärtner Willh. Krieger im hiesigen botanischen Garten an einem ganz ins Freie (ohne Topf) und in sonniger Lage gejiflanzten, 5—6 Jahre alten Exemplare der Aralia papyrifera Hook. eine kaum erst aus dem Boden hervorsprossende, gelblichte Schmarotzerpflanze, die er für eine Orobanche hielt und als der Aralia schädlich, wegschnitt und fortwarf, worauf sie sich an diesem Exemplar nicht wieder zeigte. ... Crescit in rad. Araliae papyriferae Hook. [Aralia papyrifera Hook.; Tetrapanax papyrifer (Hook.) K. Koch; Fatsia papyrifera (Hook.) Miq. ex Witte (Araliaceae)]" = Orobanche stenantha Lojac. in Naturalista Sicil. 2(3): 62 (1882) (Beck 1930: 219). Syn: ≡ Orobanche hederae f. stenantha (Lojac.) Beck in Biblioth. Bot. 19: 260 (1890). Ind.loc.: Italy: "In fruticetis submontanis sylvaticis ad radices... in Val Demone [Sicily]. Junio." = Orobanche boromaea Rion ex Beck in Biblioth. Bot. 19: 260 (1890) ["Orobanche Boromaea Rion, Exsicc. Vidi spec. orig."], pro syn. = Orobanche ciliata Zahlbr. ex Beck in Biblioth. Bot. 19: 260 (1890) ["Orobanche ciliata Zahlbr. msc. in herb. Univ. Pragensis"], pro syn. Syn: = Orobanche stenantha Lojac. in Naturalista Sicil. 2(3): 62 (1882) (Beck 1890: 260) = Orobanche racemosa Schleich. ex Beck in Biblioth. Bot. 19: 260 (1890) ["Orobanche racemosa Schleicher, Exsicc. Vidi orig."], pro syn. = Orobanche balearica Sennen & Pau in Butll. Inst. Catalana Hist. Nat. 8: 18 (1911) ["13. — Orobanche (Phaelipea) balearica Sennen et Pau."] (Beck 1930: 219). Ind.loc.: Spain, Balearic Islands: "Mis amigos, los Hermanos Sennen y Elías, han tenido la bondad de comunicarme unas cuantas plantas, recogidas en las cercanías de Sóller [Mallorca]" (Pau 1911: 18). Herbarium Specimens: Syntypes: "Plantes d'Espagne — F. Sennen / N.º 2018 / Orobanche balearica Sen et Pau / Baléares: Alfabia, près de Palma, / sur le lierre / 1914-18-VI Leg. F. Bianor" (BR 0000026151717V - BR0000026151717V.jpg - https://www.gbif.org/es/occurrence/1840543035, rev. Ó. Sánchez Pedraja Jul 2020; https://data.biodiversitydata.nl/naturalis/specimen/L.2817406 - L.2817406.jpg, rev. Ó. Sánchez Pedraja Jul 2020; idem (US 2128316 - Image No. Barcode 03926915 - http://n2t.net/ark:/65665/m34446de17-3a51-4ff3-a054-0ae9d91a2950 - https://www.gbif.org/es/occurrence/2625843870, rev. Ó. Sánchez Pedraja Jun 2022); "N.º 2018 / Orobanche balearica Sen et Pau / [Baléares: Alfabia, près de Palma, sur le lierre] 1914-18-VI / Leg. F. Bianor" (LY 0475996 - hb. R. Bonaparte - https://mediaphoto.mnhn.fr/media/15451552944546CuT76I8Atb1qOmm - https://explore.recolnat.org/occurrence/2FF9139E841B4110BAD5CD0BBC9B5775, rev. Ó. Sánchez Pedraja Jun 2022). Obs. PI. d'Esp. n. 2018 = Orobanche minor var. "β" Bertol., Fl. Ital. 6: 421[-422] (1846) ["Varietatem β"], nom. illeg. (Hausmann 1852: 651). Ind. loc.: Italy: "Varietatem β legi Sarzanae prope arcem di Sarzanello [La Spezia, Liguria], et in Montedarme supra Hederam Helicem L. Accepi Genuâ [Genoa / Genova, Liguria] pariter lectam supra Hederam Helicem L. in villa Eq. De Nigro a Rev. Vauchero, et a Rainero [Piedmont], Neapoli [Campania] in sylva di Portici, et a Capo di monte e radicibus Hederae Helicis L." = Orobanche hederae var. microcalyx Hausm. ex Rchb. f., Icon. Fl. Germ. Helv. 20: 103, t. 216, f. 3 (1862) ["b. microcalyx v Hausm. in litt."]. Ind. loc.: Germany: "b. microcalyx: häufig um Botzen [Bötzen, Jesewitz, Nordsachsen, Saxony] an Abhängeu [Weinbergen, Unstrut-Hainich, Thuringia], in sonnigen Lagen auschliesslich auf Hedera Helix schmarotzend. v. Hausmann! qui ab anno 1844. eam observavit". Obs. "Caulis flavidus" ? = Orobanche amethystea var. hyrcana Beck in Biblioth. Bot. 19: 231 (1890). ["ε (Orobanche) hyrcana"] (Schiman-Czeika 1964: 20). Ind. loc.: Iran: "Persia borealis: prope Asterabad [Gorgan / Esterabad, Golestan Province] (herb. Fischer, Petrop.)". Obs. "stigma luteum" (Beck 1890: 231) = Orobanche hederae var. trichodea J.A. Guim. in Brotéria 3: 181 (1904) ["β. (O.) trichodea Guim."]. Ind. loc.: Portugal: "solum (O.) trichodea Guim., in Lusitania boreali pervulgata, in provinciis australibus nunquam est inventa ... Alemdouro littoral:—Valença: [Viana do Castelo District] Costa da E'rvilha (R. da Cunha!); Espinhal [Coimbra District] (A.J. Freiré!). Centro littoral:—Arredores de Coimbra (A. de Carvalho!), Santa Cruz (Lopes Manita!), cerca do Jardim Botánico (Loureiro!), muro da Penitenciaria (M. de Lucena!)." = Orobanche hederae f. gyroflexa Beck in Biblioth. Bot. 19: 260 (1890) ["Formas...: 3. (Orobanche) gyroflexa"] (Beck 1890: 260, "Corollae acriter pronus curvatae; linea dorsalis semicirculata. Spica nunc densi- nunc laxiflora. In ceteris typo aequalis") = Orobanche hederae f. megaphyllon Beck in Biblioth. Bot. 19: 261 (1890) ["Formas...: 6. (Orobanche) megaphyllon"] (Beck 1890: 261, "Bracteae corollas conspicue superantes. Onines varietates cum typica occursant.") = Orobanche hederae f. minuscula Beck in Biblioth. Bot. 19: 261 (1890) ["Formas...: 5. (Orobanche) minuscula"] (Beck 1890: 261, "Corolla minima —1 cm longa.") = Orobanche hederae f. monochroa Beck in Biblioth. Bot. 19: 260 (1890) ["Formas...: 4. (Orobanche) monochroa"] (Beck 1890: 260, "Corolla tota albo-lutea") = Orobanche hederae f. stenantha (Lojac.) Beck in Biblioth. Bot. 19: 260 (1890) ["Formas...: 2. (Orobanche) stenantha Lo Jacono"]. Syn: ≡ Orobanche stenantha Lojac. in Naturalista Sicil. 2(3): 62 (1882) =
Orobanche
hederae f. godronii
Rouy in Rouy & Foucaud, Fl.
France 11: 190 (1909) (Beck
1930:
222).
Syn: =
Orobanche laurina sensu Godron,
non Bonap. ex Bertol. Fl. Ital. 6: 424 (1846).
Herbarium Specimens: 1.
"Orobanche laurina Godr.! an Bert. / Jardin du plantes 4 juillet 1804
/ Sur le lierre! ..." – 2. "Orobanche
= Orobanche hederae f. immaniflora Beck in Repert. Spec. Nov. Regni Veg. 18: 40[472] (1922) ["Forma 3. immaniflora G. Beck"]. Ind. loc.: Germany: "Germania: ad arcem Liebenstein [Thuringia] prope Kamp ad fl. Rhenum (Spiessen) in exemplaribus pluribus." Misapplied names: – Orobanche major sensu L. in Loefl., Iter Hispan.: 151 (1758), non Orobanche major L., Sp. Pl.: 632 (1753), nom. rej. [= Orobanche caryophyllacea Sm. (1798)] (Reuter 1847: 29, sub O. Hederae Duby). Herbarium Specimens: 1. "429 / Orobanche Hederae Vaucher! / Parasite sur le liérre / Vallée de Djimil [40°53'58.0"N 41°07'58.4"E, Kaçkar Mountains / Lazistan Mountains, Pontic Mountains / Kuzey Anadolu Dağları] (Lazistan), vers / 2000 mètres d'altitude / Juillet 1866 [B. Balansa, Plantes d'Orient, 1866?]". – 2. "Orobanche Hederae Duby Reut. / in DC. prod. / O. major Loeflin! non Linn. [m. Reuter]" (G 00769188 - n° SIB 436398/1 - https://www.ville-ge.ch/musinfo/bd/cjb/chg/adetail.php?id=559080 - https://www.ville-ge.ch/imagezoom/?fif=cjbiip/cjb56/img_20170215/G00769188.ptif&cvt=jpeg; rev. Ó. Sánchez Pedraja Jun 2022). – Orobanche barbata sensu Rchb., Fl. Germ. Excurs. 1 (1831-32) [saltem p.p.], non Orobanche barbata Poir. in Lam., Encycl. 4: 621 (1797). Ind. loc.: "An Hedera Helix im südlilichsten Gebiete, in Piemont [Piedmont / Piemonte; Italy] ... Gênes nach Savone [Liguria, Italy] ... Ponente ... bei Gênes [Genoa] dans le jardins Négri: Vaucher". Obs. "stigma flavum" – Orobanche minor sensu Benth., Cat. Pl. Pyrénées: 109 (1826) (p.p. ["Elle croît sur ... le Hedera Helix"]), non Orobanche minor Sm., Engl. Bot. 6: 422 (1796-1797) – Orobanche medicaginis sensu Rchb., Iconogr. Bot. Pl. Crit. 7: 41, t. 684 f. 919 (1829), non Orobanche medicaginis Duby, Bot. Gall.: 349 (1828) [= O. lutea Baumg. (1816)]. Syn.: = Orobanche du Lierre Vaucher, Monogr. Orobanch.: 56, t. 8 (1827), nom. illeg. (Reichenbach 1829: 42; Reichenbach fil. 1862: 103, sub O. Hederae Duby). Ind. loc.: "Habitat in muris arcis Genevensis [Geneve / Genève, Switzerland] ... Vaucher; in muro vetusto ad viam regiam prope Viviscum [Vevey, Vaud / Waadt, Switzerland] Gaudin". Obs. "Corollis ... fauce constrictis" (Reichenbach 1829: 42; Grisebach 1844: 57; Reichenbach fil. 1862: 102) |
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Ind. loc.: "Hederae parasiticam in Vasconiâ [Vasconia, Spain-France] reperiit cl. Des Moul. (v.s. ex Genuâ.)" (Duby 1828: 350). "Cette singulière Orobanche se trouve sur les collines qui s'étendent de Gênes à Savone [Genoa / Genova to Savone / Savona, Liguria, Italy]. Je l'ai cueillie en 1825, sur les hauteurs qui dominent cette dernière ville, et dans les maisons de campagne du Ponente. Je l'ai revue l'année suivante en grande abondance à Gênes même, dans les jardins Négri. Elle vient d'être retrouvée par M. Charles Des Moulins, dans le Périgord [Dordogne, Nouvelle-Aquitaine, France] et aux environs de Libourne [Gironde, Nouvelle-Aquitaine, France]. Elle fleurit dans le mois de Juin." (Vaucher 1827: 57). Typus / Type: "Type in Berne" (Novopokrovskij & Tzvelev 1958: 98[104], sub O. hederae Duby). "(v.s. ex Genuâ [G]) (Duby 1828: 350)" (Gilli 1982: 19; Piwowarczyk & al. 2023: 71, sub O. hederae Duby). Designated by: "holotype indicated by (Gilli 1982: 19)" (Piwowarczyk & al. 2023: 71, sub O. hederae Duby). Host: Predominantly on Araliaceae species, usually on Hedera sp. (mainly on H. helix L. [aggr.]), more rarely on other genera Fatsia, × Fatshedera, Tetrapanax and Kalopanax (Rumsey & Jury 1991: 273, sub O. hederae Duby) and also mentioned on Geraniaceae. Araliaceae Juss. Eleutherococcus wardii (W.W. Sm.) S.Y. Hu [≡ Acanthopanax wardii W.W. Sm.]. Garden weed. Europe: Botan. Garten Dresden (Uhlich & al. 1995: 116, sub O. hederae). Fatsia japonica (Thunb.) Decne. & Planch. [≡ Aralia japonica Thunb.]. Garden weed. Europe: France and Portugal (P 00598694.jpg; Guimarães 1904: 183; Beck 1930: 220). Hedera algeriensis Hibberd [= Hedera canariensis f. variegata (Paul) Verhaeghe]. Tunisia (El Mokni & al. 2015: 8, sub O. hederae). Hedera canariensis Willd. [≡ Hedera helix var. canariensis (Willd.) DC.]. Canary Islands and Morocco (Jahandiez & Maire 1934: 699, sub O. Hederae Duby). Hedera colchica (K. Koch) K. Koch [≡ Hedera helix var. colchica K. Koch]. Caucasus (Beck 1930: 220, "f. Woronow et Bolle"; Piwowarczyk & Tatanov 2013: 26, sub O. hederae; Piwowarczyk & al. 2023: 71, sub O. hederae Duby). Hedera helix L. Europe and Caucasus (Beck 1890: 261; Beck 1930: 220; Maslo 2016: 67, sub O. hederae; Piwowarczyk & al. 2023: 71, sub O. hederae Duby). Hedera hibernica Poit. [≡ Hedera helix var. hibernica (Poit.) G. Kirchn.]. Atlantic Europe (Rumsey & Jury 1991: 273, sub O. hederae Duby). Hedera taurica (Hibberd) Carrière [≡ Hedera helix var. taurica Hibberd]. Krim / Crimea / Крым / Крим / Tauric Peninsula (Tsymbalyuk & Mosyakin 2013: 724, sub O. hederae Duby). Kalopanax septemlobus (Thunb.) Koidz. [≡ Acanthopanax septemlobus (Thunb.) Koidz. ex Rehder]. Garden weed. Europe: Botan. Garten Dresden (Uhlich & al. 1995: 116, sub O. hederae). Tetrapanax papyrifer (Hook.) K. Koch [≡ Araliae papyrifera Hook.; ≡ Fatsia papyrifera (Hook.) Miq. ex Witte]. Garden weed (Meissner 1866: 17, sub O. aralioctona; Beck 1890: 261; Beck 1930: 219; Beck 1930: 220). Geraniaceae Juss. Geranium reuteri Aedo & Muñoz Garm. [= Geranium canariense Reut., nom. illeg.]. Canary Islands [Tenerife Island] (Gilli 1980: 115, sub O. hederae var. microcalyx). Comments: Chorology obs.: "In some places, it represents a plague on ornamental ivies" (Piwowarczyk & al. 2023: 71, sub O. hederae Duby). Phylogenetic obs.: Carlón & al. in Documentos Jard. Bot. Atlántico (Gijón) 1: 25, 26-27 (2002) et Documentos Jard. Bot. Atlántico (Gijón) 3: 61 f. 4 ["Phylogenetic relationships"], 64 (2005). Chromosomatic number: n = 19 (Palomeque & Sañudo 1981 "Jaén, Spain"; Fernández López & Palomeque 1984 "Jaén, Spain"); 2n = 38 (Weber 1976; Schneeweiss & al. 2004). Ic.: a, inflorescences (on Hedera helix L.; Ontón, Spain); b, flowers (id.); c, flower structure (on Hedera sp.; Santander, Spain); d, flower section (id.); e, Beck 1890: t. 4 f. 83; f, Hauts-de-Seine, France (P 00598694.jpg); Georgia (Piwowarczyk & al. 2023: 72 f. 46, sub O. hederae Duby). Distribution: Africa [native (Algeria, Macaronesia [Azores and Canary Islands], Morocco and Tunisia) and naturalized / garden weed (South Africa)]. America [naturalized / garden weed (USA)]. Asia [native (Caucasus [Azerbaijan, Georgia and Russia], Iran, Syria and Turkey) and naturalized / garden weed (Philippines)]. Europe [native and naturalized / garden weed (Albania, Austria [naturalized], Azerbaijan, Balearic Islands, Belgium, Bosnia and Herzegovina, Bulgaria, Corsica, Croatia, Czech Republic [naturalized], Cyprus [unconfirmed], Denmark [naturalized], England [United Kingdom], France, Georgia, Germany [native and naturalized / garden weed], Greece, Hungary [naturalized, needs be confirmed], Ireland, Italy, Krim, Liechtenstein, Luxembourg, Malta, Montenegro, Netherlands, North Macedonia, Norway [naturalized], Poland [naturalized], Portugal, Romania, Russia, Scotland [United Kingdom] [naturalized], Serbia [needs be confirmed], Slovakia [unconfirmed], Slovenia, Spain, Sweden [naturalized], Switzerland, Turkey, Ukraine [alien] and Wales [United Kingdom)]. Global distribution (GBIF; accessed Apr 2014); Distributional range (GRIN; accessed Apr 2014); https://www.discoverlife.org...hederae&guide=North_American_Invasives&flags=HAS: (Plants Database, United States Department of Agriculture, accessed Jun 2022); https://www.cabi.org/isc/datasheet/116001 (Cabi Jun 2022). |
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Specimens examined and literature records. Africa / África / Afrique / Afrika / Африка / 非洲 (Beck 1890: 261, sub O. Hederae Duby; Beck 1930: 221, sub O. Hederae Duby; Pusch 2009: 64 Abb. 80, sub O. hederae Duby; El Mokni & al. 2015, sub O. hederae) Algeria / الجزائر / Argelia / Algérie / Algerien (Beck 1890: 261, sub O. Hederae Duby; Jahandiez & Maire 1934: 699, sub O. Hederae Duby; Pusch 2009: 64 Abb. 80, sub O. hederae Duby) Aïn Defla Province: 1. "Université d Alger / Herbier de l'Afrique du Nord / Orobanche Hederae Duby. / A. Miliana, sur Hedera Helix dans / les travertins / 12.7 1917 Dr. R. Maire" (https://science.mnhn.fr/institution/um/collection/mpu/item/mpu261524 - MPU261524.jpg - hb. Maire - http://www.gbif.org/occurrence/1424915976 - https://explore.recolnat.org/occurrence/4BE733DAB904483099F17564593CC551; rev. Ó. Sánchez Pedraja Jul 2020). Algiers Province: "Algeria: La caldo" (Beck 1890: 261, sub O. Hederae Duby); "Algier, bei La Caldo!" (Beck 1930: 221, sub O. Hederae Duby). Constantine Province: "Constantine [36°25'05.8"N 6°38'35.0"E], Algérie - 13 May 2020" (https://www.inaturalist.org/observations/46151390 by Karim Haddad in iNaturalist; accessed Jul 2020). Tlemcen Wilaya: "in the Mountains of Tlemcen belonging to O3 and more precisely at the site of the Hafir station at elevation of 1200 m, 26 May 2010, collected by Babali, (ENSA13427)" (Aissaoui & al. 2023: 14, sub O. hederae Duby). Macaronesia / Macaronésie / Makaronesien Azores / Açores (Portugal) (http://www.azoresbioportal.angra.uac.pt...id=F00706; accessed Jul 2020) Canarias, Islas / Canary Islands / Kanarische Inseln (Spain / España / Espagne / Spanien) (Gilli 1980: 115; Gilli 1980: 115, sub O. hederae var. microcalyx; Reyes, pers. comm., Jul 2020) Santa Cruz de Tenerife / Santa Cruz Province La Gomera: "El Cedro [28°07'31.0"N 17°13'24.1"W 28.125287, -17.223347], 3 May 2023" (https://www.inaturalist.org/observations/159501865 by Audrey Ipalan Gomera in iNaturalist; accessed Aug 2023 [specimens in the preanthesis, without determining host plant, need confirmation]). La Palma / San Miguel de La Palma Island: "O. hederae var. hederae: La Palma, Ostseite, Waldgebiet Los Tilos bei Los Sauces, 400 m, 13.7.1977" (Gilli 1980: 115); "La Palma [28°47'08.5"N 17°48'23.0"W 28.785700, -17.806400, Los Sauces, San Andrés y Sauces], 2014-06-25" (https://observation.org/observation/86326843 - https://www.gbif.org/occurrence/2832528067 by Mark Broeckaert in Observation.org; accessed Aug 2023 [typical form, apparently on Hedera sp.]); "La Palma, San Andrés y Sauces [28°47'04.3"N 17°48'27.6"W], Los Tilos [Los Tiles] (682 m), 27 Aug 2018" (https://www.inaturalist.org/observations/45247417 - https://www.gbif.org/occurrence/2626103012 by Sylvain Piry in iNaturalist; accessed Jul 2020 [typical form, apparently on Hedera sp.]); "Puntallana [28°45'40.3"N 17°46'46.6"W 28.761206, -17.779613, Llano Molino], 8 Jul 2021" (https://www.inaturalist.org/observations/87154188 by https://www.inaturalist.org/people/srrobin in iNaturalist; accessed Aug 2023 [yellow form]); "Parque Natural Las Nieves, San Andrés y Sauces [28°46'50.6"N 17°48'33.2"W 28.780725, -17.809222], 7 Ago 2022" (https://www.inaturalist.org/observations/130399627 - https://www.gbif.org/occurrence/3892300721 by Oriol Sastre; accessed Aug 2023 [typical form, apparently on Hedera sp.]); "Parque Natural Las Nieves, San Andrés y Sauces [28°46'38.0"N 17°48'59.3"W 28.777217, -17.816475], 31 Ago 2022" (https://www.inaturalist.org/observations/133587136 by Emilio Valbuena-Ureña in iNaturalist; accessed Aug 2023 [apparently on Hedera sp.]); "38729, Santa Cruz de Ténérife [28°47'15.6"N 17°48'21.6"W 28.787664, -17.805997, San Andrés y Sauces], 29 May 2023" (https://www.inaturalist.org/observations/164531371 by https://www.inaturalist.org/people/qfaye in iNaturalist; accessed Aug 2023 [apparently on Hedera sp.]). Tenerife Island: "O. hederae var. microcalyx: Tenerife, Teno-Gebirge [Rural de Teno Park] westlich Erjos, Lorbeerwad, auf Geranium canariense [this species is not a host-plant of O. hederae], 6.3.1977" (Gilli 1980: 115, sub O. hederae var. microcalyx); "Monte del Agua [28°19'29.4"N 16°48'48.3"W 28.324826, -16.813411, Los Silos (Erjos)], ~1000 m, 23 Jun 2021" (https://www.inaturalist.org/observations/84284680 - https://www.gbif.org/occurrence/3314438460 by Michael 2020 [https://www.inaturalist.org/people/miwi2020] in iNaturalist; accessed Aug 2023 [yellowish form with pale yellow stigma, on Geranium cf. reuteri Aedo & Muñoz Garm.]); "Santa Cruz de Tenerife [28°19'56.8"N 16°49'39.6"W 28.332447, -16.827677, Los Silos, Las Lagunetas (Buenavista del Norte)], 28 Jun 2021 / The host is probably Geranium reuteri Aedo & Muñoz Garm. [= Geranium canariense Reut., nom. illeg.]" (https://www.inaturalist.org/observations/84950153 - https://www.gbif.org/occurrence/3327735549 by https://www.inaturalist.org/people/hchavez in iNaturalist; accessed Aug 2023 [yellowish form with pale yellow stigma]); "Almáciga, 38129 [28°34'16.7"N 16°10'07.1"W 28.571316, -16.168649, El Draguillo (Distrito de Anaga)], 11 Feb 2023" (https://www.inaturalist.org/observations/149067380 by David García del Olmo in iNaturalist; accessed Aug 2023 [on Hederae sp.]); "Santa Cruz de Tenerife [Los Silos, 28°19'17.3"N 16°49'18.0"W 28.321472, -16.821667 (Erjos)], 14 May 2023" (https://www.inaturalist.org/observations/161690774 - https://www.gbif.org/occurrence/4116166754 by Adrián Martínez Martínez in iNaturalist; accessed Aug 2023 [yellowish form with pale yellow stigma, on Geranium cf. canariense?]); "38479 Los Silos [28°20'19.8"N 16°49'59.5"W 28.338823, -16.833183, Las Lagunetas (Buenavista del Norte)], 2 Jun 2023" (https://www.inaturalist.org/observations/165976882 by https://www.inaturalist.org/people/qfaye in iNaturalist; accessed Aug 2023 [yellowish form with pale yellow stigma, on Geranium cf. reuteri Aedo & Muñoz Garm. [= Geranium canariense Reut., nom. illeg.]). Morocco / المغرب, / Marruecos / Marokko (Jahandiez & Maire 1934: 699, sub O. Hederae Duby; Sennen & Mauricio 1934: 87, sub O. Hederae L.; Emberger & Maire 1941: 1130, sub O. Hederae Duby; Dobignard & Jordan 1987: 94; Rumsey & Jury 2002: 592, sub O. hederae Duby; Pusch 2009: 64 Abb. 80, sub O. hederae Duby; Fennane 2018: 14, sub O. hederae Duby) "HA-4 (Mgoun à Akka-n-Tazaght) [Dobignard & Jordan 1987: 94]" (Fennane 2018: 14, sub O. hederae Duby). Ifrane Province: "Sur le Lierre (Hederá Helix L. ssp. canariensis (Willd.) Maire [Hedera canariensis Willd.] ... MA. Ifrane (Emberger)" (Jahandiez & Maire 1934: 699, sub O. Hederae Duby); "environ 19 km d'Azrou vers Midelt [Valdés 2013:87])" (Fennane 2018: 14, sub O. hederae Duby). Nador Province: Gurugú (Sennen & Mauricio 1934: 87, sub O. Hederae L.); "R. Mont Gourougou [Gurugú] (S. et Ma. [Sennen et Mauricio])" (Emberger & Maire 1941: 1130, sub O. Hederae Duby); Gareb [Garet 34°57'36.0"N 3°06'00.0"W] (Rumsey & Jury 2002: 592, sub O. hederae Duby). South Africa / Suid-Afrika / Sudáfrica / Afrique du Sud / Südafrika [naturalized / garden weed] "Cape Winelands [33°56'07.2"S 18°52'31.8"E -33.935337, 18.875500, Stellenbosch], ZA-WC, ZA, 3 Nov 2019" (https://www.inaturalist.org/observations/35240740 - https://www.gbif.org/occurrence/2529347161 by Quentin Groom in iNaturalist; accessed Jul 2020); "Stellenbosch University [33°55'53.4"S 18°51'56.8"E -33.931512, 18.865770], Stellenbosch, WC, ZA, 17 Nov 2021" (https://www.inaturalist.org/observations/101319853 by Dave Richardson in iNaturalist; accessed Aug 2023). Tunisia / تونس / Tunisie / Túnez / Tunisien (El Mokni & al. 2015: 8, sub O. hederae) Jendouba Governorate: "Aîn Draham (Jendouba), 36°45′23" N, 08°41′32" E, 700 m a.s.l., on Hedera algeriensis Hibberd, 03.7.2011, R. El Mokni (BIZ!)." (El Mokni & al. 2015: 8, sub O. hederae). America / América / Amérique / Amerika / Америка / 美洲 [naturalized / garden weed] United States of America (USA) / Estados Unidos de América / États-Unis d'Amérique / Vereinigte Staaten von Amerika / Соединенные Штаты Америки [naturalized / garden weed] California: "Alameda Co.: UC-Berkeley campus, small persisting colony in Hedera groundcover next to Koshland Hall. June 7, 2000, B. Ertter 17310 (UC); between student center and Alumni House. May 8, 2001, B. Ertter 17626 (UC)" (Hrusa & al. 2002; O. hederae in Jepson Flora Project [County distribution, Specimen records, Consortium of California Herbaria]; accessed Apr 2014); "UC Berkeley campus, west end University Drive (Alameda County, California, US) / May 28, 2011" (http://calphotos.berkeley.edu/cgi/img_query?enlarge=0000+0000+0911+1524 by Neal Kramer in CalPhotos Photo Database; accessed Apr 2014); "Orobanche minor, growing with invasive ivy [Hedera sp.?], University of California at Berkeley Campus, Alameda Co., CA, 14 Aug 2012." (photos 1, 2, 3 by Mark Egger, accessed Feb 2014 [these photos seem to correspond to O. hederae Duby], accessed et rev. Ó. Sánchez Pedraja Feb 2014); "University of California - Berkeley [37°52'19.3"N 122°15'55.1"W 37.872017, -122.265312], 15 Jul 2023" (https://www.inaturalist.org/observations/173162068 - https://www.gbif.org/occurrence/4162352839 by Arvel Hernandez in iNaturalist; accessed Aug 2023 [reddish form]). Asia / Asie / Asien / Азия / 亚洲 (Beck 1890: 261, O. Hederae Duby; Beck 1930: 221, sub O. Hederae Duby; Novopokrovskij & Tzvelev 1958: 98[104], sub O. hederae Duby; Pusch 2009: 64 Abb. 80, sub O. hederae Duby) "Caucasus: All regions [Russia], besides southern Transcaucasia [see Azerbaijan, Georgia]" (Novopokrovskij & Tzvelev 1958: 98[104], sub O. hederae Duby). Iran / ایران / Irán [Persia] (Beck 1890: 231, loc. class. of O. amethystea var. hyrcana; Schiman-Czeika 1964: 18, sub O. Hederae Duby) Gilan Province: "N: Gilan: Tspili [Espeyli, Siahkal County], Linds. 1130!" (Schiman-Czeika 1964: 18, sub O. Hederae Duby). Golestan Province: "Persia borealis: prope Asterabad [Gorgan / Esterabad] (herb. Fischer, Petrop.)" (Beck 1890: 231, loc. class. of O. amethystea var. hyrcana). Mazandaran Province: "Mazandaran Province [36°27'32.9"N 51°45'52.7"E - 36.459136, 51.764648, near Kodir]", 4 Jun 2009 (https://www.inaturalist.org/observations/105665907 - https://www.gbif.org/es/occurrence/3466198810 by Shahrzad Fattahi in iNaturalist; accessed Jun 2022). Pilipinas / Filipinas / Philippines / Philippinen / Филиппины / 菲律宾 [naturalized?] Caraga Region / Region XIII: Surigao del Sur (https://www.gbif.org/es/occurrence/1265649891 [n.v.]). Syria / Siria / Syrien / Syrie Latakia / Ladhakia Governorate: "Latakia [35°44'58.8"N 36°00'40.9"E - 35.749678, 36.011347, near Alsukkariah], 4 Jun 2021" (https://www.inaturalist.org/observations/81636120 - https://www.gbif.org/es/occurrence/3302347841 by Safwan Dawod in iNaturalist; accessed Jun 2022). Tartus Governorate / Tartous: "Ash-Shaykh Badr [35°02'20.9"N 36°03'24.3"E 35.039139, 36.056750], SY-TA, SY, 22 May 2023" (https://www.inaturalist.org/observations/163158678 by Ali Ibrahim in iNaturalist; accessed Aug 2023). Turkey / Türkiye / Turquía / Турция (Beck 1890: 261, O. Hederae Duby; Beck 1930: 221, sub O. Hederae Duby; Pils 2006: 259, sub O. hederae; Pusch 2009: 64 Abb. 80, sub O. hederae Duby) Thrace (European Turkey) Marmara Region / Marmara Bölgesi Istanbul Province (European Turkey): "Karaburun [41°20'27.2"N 28°39'51.3"E], 34277 Arnavutköy / İstanbul" (https://www.inaturalist.org/observations/46831979 by Bilâl Erdoğan in iNaturalist; accessed Jul 2020 [on Hedera helix aggr.]); "Adalar [40°50'58.7"N 29°07'09.8"E 40.849625, 29.119397], İstanbul, 25 May 2023" (https://www.inaturalist.org/observations/164800473 - https://www.gbif.org/es/occurrence/4129981747 by Filip Kawa, sub O. hederae, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Aug 2023, sub O. pubescens [yellow form]). Anatolia (Asiatic Turkey) Black Sea Region / Karadeniz Bölgesi Rize Province / Rize ili: 1. "429 / Orobanche Hederae Vaucher! / Parasite sur le liérre / Vallée de Djimil [40°53'58.0"N 41°07'58.4"E, Kaçkar Mountains / Lazistan Mountains, Pontic Mountains / Kuzey Anadolu Dağları] (Lazistan), vers / 2000 mètres d'altitude / Juillet 1866 [B. Balansa, Plantes d'Orient, 1866?]". – 2. "Orobanche Hederae Duby Reut. / in DC. prod. / O. major Loeflin! non Linn. [m. Reuter]" (G00769188 - n° SIB 436398/1 - https://www.ville-ge.ch/musinfo/bd/cjb/chg/adetail.php?id=559080 - https://www.ville-ge.ch/imagezoom/?fif=cjbiip/cjb56/img_20170215/G00769188.ptif&cvt=jpeg; rev. Ó. Sánchez Pedraja Jun 2022); "B. Balansa, Plantes d'Orient, 1866 / Orobanche Hederae Vauch. / (Reuter) / Parasite sur le liérre / Région Sous-alpine du Lazistan / près de Djimil [40°53'58.0"N 41°07'58.4"E, Kaçkar Mountains / Lazistan Mountains, Pontic Mountains / Kuzey Anadolu Dağları], vers / 2000 mètres d'altitude / Juillet" (https://science.mnhn.fr/institution/mnhn/collection/p/item/p02968125 - P02968125.jpg - hb. Drake - http://www.gbif.org/occurrence/437280446 - https://explore.recolnat.org/occurrence/0D7F0578B3234B0AA5085E5FB3E4B8D6; rev. Ó. Sánchez Pedraja Aug 2023). East Black Sea Region / Doğu Karadeniz Bölgesi Artvin Province / Artvin ili: "im Distr. Artwin bei Ardanudsh im Mai 1907 (Woronow!)" (Beck 1930: 221, sub O. Hederae Duby). Marmara Region / Marmara Bölgesi Istanbul Province (Asiatic Turkey): "Lavie Konaklari, Cekmekoy, Istanbul [41°02'49.7"N 29°10'18.5"E 41.047131, 29.171812]", 4 Jun 2021 (https://www.inaturalist.org/observations/81554540 by Cem Cizem in iNaturalist; accessed Jun 2022 [on Hedera sp.]). Kocaeli Province / Kocaeli ili: "Eşme, 41110 Kartepe/Kocaeli [40°43'54.9"N 30°10'55.9"E - 40.731909, 30.182202]", 29 May 2022 (https://www.inaturalist.org/observations/119229820 by Önder Alioğlu in iNaturalist; accessed Jun 2022). Mediterranean Region / Akdeniz Bölgesi Kahramanmaraş / Kahramanmaraş ili / Maraş Province: "in monte Tauro (Kotschy ! n. 47)" (Reuter 1847: 28, sub O. Hederae Duby); "... et in monte Tauro [Taurus Mountains / Toros Dağları] anno 1836 (leg. Kotschy)" (Beck 1890: 261, sub O. Hederae Duby). Osmaniye Province / Osmaniye ili: "Zorkun Yolu, Karacalar, 80000 Karacalar/Osmaniye Merkez [37°01'07.6"N 36°16'18.5"E 37.018783, 36.271795], 4 Jun 2017" (https://www.inaturalist.org/observations/150677886 by Mehmet Çelik in iNaturalist; accessed Aug 2023). West Black Sea Region / Batı Karadeniz Bölgesi Tokat Province / Tokat ili: "Asia minor: prope Tokat (leg. Wiedemann) ..." (Beck 1890: 261, sub O. Hederae Duby). Europe / Europa / Европа / 欧洲 (Vaucher 1827: 57, sub Orobanche du Lierre; Grenier 1853: 640, sub O. Hederae (O. du Lierre) Vauch.; Reichenbach fil. 1862: 104, t. 181, sub O. laurina; Beck 1890: 261, sub O. Hederae Duby; Beck 1930: 220-221, sub O. Hederae Duby; Pusch 2009: 64 Abb. 80, sub O. hederae Duby) Albania / Shqipëri / Shqipëria / Shqipnia (Barina & al. 2011: 335, sub O. hederae Duby) Fier District / Rrethi i Fierit: "loc. 7 [in Kurora natura reserve next to village Adriatik (Alibejas), in softwood forest, 40.8118°N, 19.5077°E, 10 m, 11.8.2010, Z. Barina & D. Pifkó]: 18174" (Barina & al. 2011: 335, sub O. hederae Duby). Vlorë County / Qarku i Vlorës: "Shëndëlli [39°44'01.8"N 20°01'45.5"E 39.733825, 20.029291], 8 May 2023" (https://www.inaturalist.org/observations/161477717 - https://www.gbif.org/es/occurrence/4116316348 by https://www.inaturalist.org/people/claudio2claudio in iNaturalist; accessed Aug 2023 [yellow form]). Austria / Österreich / Autriche [naturalized] (Beck 1890: 261, sub O. Hederae Duby; Beck 1930: 221, sub O. Hederae Duby; Pusch 2009: 64 Abb. 80, sub O. hederae Duby) Lower Austria / Niederösterreich: "2500 Baden [Baden bei Wien, 48°00'38.9"N 16°13'36.4"E 48.010812, 16.226765], 24 Jun 2023" (https://www.inaturalist.org/observations/169208685 by Michael Andresek in iNaturalist; accessed Aug 2023). Vienna Province: "Austria: in horto botanico Vindobonensi [Botanical Garden of the University of Vienna, Vienna / Wien] certe introducta" (Beck 1890: 261, sub O. Hederae Duby); "1030 Wien [48°11'29.4"N 16°22'59.8"E 48.191505, 16.383282], 21 Jun 2023" (https://www.inaturalist.org/observations/168711637 by Martin Scheuch in iNaturalist; accessed Aug 2023). Azerbaijan / Azərbaycan / Азәрбајҹан / Azerbaiyán / Aserbaidschan / Azerbaïdjan (Beck 1930: 221, sub O. Hederae Duby; Karjagin 1957; Pusch 2009: 64 Abb. 80, sub O. hederae Duby) Lənkəran / Lankaran District / Ленкоранский район: "Prov. Baku, bei Lenkoran [Lənkəran / Lankaran] im Mai (Radde!)" (Beck 1930: 221, sub O. Hederae Duby). Belgique / Belgium / België / Bélgica (Beck 1890: 261, sub O. Hederae Duby; Beck 1930: 220, sub O. Hederae Duby; Pusch 2009: 64 Abb. 80, sub O. hederae Duby) Flemish Brabant / Vlaams-Brabant / Brabante Flamenco: "Belgia: ... Tierlemont [Tienen]" (Beck 1890: 261, sub O. Hederae Duby). Namur Province / Nameur / Namen: "Belgia: Namur ..." (Beck 1890: 261, sub O. Hederae Duby). Bosnia and Herzegovina / Bosnia y Herzegovina / Bosna i Hercegovina / Босна и Херцеговина (Maslo 2014: 135, sub O. hederae; Maslo 2016: 67, sub O. hederae) Herzegovina-Neretva Canton / Hercegovačko-neretvanski kanton 7 Херцеговачко-неретвански кантон / Hercegovačko-neretvanska županija: "Two localities of O. hederae have been recorded during the field investigations in the area of Mostar (Maslo, 2014). The first finding was recorded on the left bank of the Neretva River just north of Carinski bridge and a small population on the right bank of the river Radobolja in the city quarter Podhum. Another small population was noted in the town of Blagaj, at the source of the Buna River (Maslo & Abadžić, 2015). Also found at sandy banks of the River Trebižat, ca 50 m downstream from the waterfall Kravice, on the right bank of the river. A few plants were found beneath Alnus glutinosa (L.) Gaertner, sheltered by bushes, parasitic on Hedera helix." (Maslo 2016: 67, sub O. hederae). Bulgaria / България / Bulgarie / Bulgarien (Pusch 2009: 64 Abb. 80, sub O. hederae Duby; Stoyanov 2013: 288, sub O. hederae Duby) Plovdiv Province / Област Пловдив: "Rhodopi Mts. (Central): along river Slivodolska, Chervenata Stena Reserve, 435 m, 41°55'17.0"N 24°50'06.0"E [41.921389, 24.835000], LG24, parasiting on the roots of Hedera helix, 04.07.2013, coll. K. Stoyanov & Ts. Raycheva (SOA 059771). A new species to Bulgaria..." (Stoyanov 2013: 288, sub O. hederae Duby). Croatia / Hrvatska Croacia / Kroatien / Croatie (Reichenbach f. 1862: 103, sub O. Hederae Duby; Beck 1890: 261, sub O. Hederae Duby; Beck 1930: 221, sub O. Hederae Duby; Pusch 2009: 64 Abb. 80, sub O. hederae Duby; Nikolić 2015) Dubrovnik-Neretva County: "Dalmatia: Ragusa [Dubrovnik], ins. Giupana [Giuppana / Šipan (Elaphiti Islands / Elafitski otoci)]" (Beck 1890: 261, sub O. Hederae Duby). Primorje-Gorski Kotar County / Primorsko-goranska županija: "Flora of Croatia / Orobanche hederae Vaucher ex Duby / Primorsko-Goranska, Otok Krk. Otok Košljun in / Puntarska Draga bay / 45°01'35.0"N 14°37'08.0"E 45.026390, 14.618890 / Understorey of forest with Pinus halepensis and Quercus ilex. / Parasitizing on Hedera helix / Leg.: Reich, D. & Hofbauer, M. / Det. M. Hofbauer & D. Reich / Field trip with P. Bildstein ..." (https://www.jacq.org/detail.php?ID=1107293 - WU 0091231 - https://wu.jacq.org/WU0091231; rev. Ó. Sánchez Pedraja Aug 2023). Šibensko-kninska županija: "Krka - donji tok [43°48'32.8"N 15°57'34.3"E 43.809108, 15.959537], Dubravice, 15, HR, 26 May 2023" (https://www.inaturalist.org/observations/163977412 - https://www.gbif.org/es/occurrence/4127045426 by https://www.inaturalist.org/people/dervogelmann in iNaturalist; accessed Aug 2023). Istria County / Istarska županija / Regione istriana: "Istria: Tribano [Triban], Momiano [Momjan] ...; Dalmatia: Ragusa, ins. Giupana" (Beck 1890: 261, sub O. Hederae Duby). Czech Republic / República Checa / Česká republika [naturalized] (Pusch 2009: 64 Abb. 80, "synanthrop", sub O. hederae Duby) Moravian-Silesian Region / Moravskoslezský kraj: "Štramberk [49°34'49.7"N 18°07'52.3"E 49.580479, 18.131203], 6 Jul 2022" (https://www.inaturalist.org/observations/124993727 by Jan Hrdý in iNaturalist; accessed Aug 2023). Cyprus / Κύπρος / Kıbrıs / Chipre / Chypre / Zypern [unconfirmed] Denmark / Danmark / Dinamarca / Danemark / Dänemark [naturalized] (Pusch 2009: 64 Abb. 80, "synanthrop", sub O. hederae Duby) Zealand / Sjælland / Själland (Pusch 2009: 64 Abb. 80, sub O. hederae Duby) Capital Region of Denmark / Region Hovedstaden: "Sjælland, København, Hovedstaden [55°39'40.5"N 12°31'37.4"E]" (https://www.inaturalist.org/observations/52348998 by persplanter in iNaturalist; accessed Jul 2020). France / Francia / Frankreich (Vaucher 1827: 57, sub Orobanche du Lierre; Duby 1828: 350, loc. class. of O. hederae; Des Moulins 1835: 80, loc. class. of O. hederae; Reuter 1847: 28, sub O. Hederae Duby; Grenier 1853: 640, sub O. Hederae (O. du Lierre) Vauch.; Reichenbach fil. 1862: 104, t. 181, sub O. laurina; Beck 1890: 261, sub O. Hederae Duby; Beck 1930: 220, sub O. Hederae Duby; Pusch 2009: 64 Abb. 80, sub O. hederae Duby) Côtes-d'Armor (22): "Orobanche hederae Vaucher ... env. Kercoz / presqu'Île Plestin / (C. du N.) / 8 août 1922" (http://coldb.mnhn.fr/catalognumber/mnhn/p/p02588155 - https://science.mnhn.fr/institution/mnhn/collection/p/item/p02588155 - P 02588155.jpg - hb. M. Paul Jovet - http://www.gbif.org/occurrence/465147419 - https://explore.recolnat.org/occurrence/5A524688C76D42E0B74760A029376592; rev. Ó. Sánchez Pedraja Jan 2014). Corsica / Corse / Corsica / Córcega (Beck 1930: 221, sub O. Hederae Duby; Habashi & Jeanmonod 2008: 42-43, sub O. hederae Vaucher ex Duby) Corse-du-Sud (2A): "Herbier E.G. Camus / 3150 / Orobanche Crithmi / Orobanche crinita Viv.? / Corse: env. de Bonifacio / 7 avril 1902 / Legit Stefani / E.G. Camus" (http://coldb.mnhn.fr/catalognumber/mnhn/p/p06903078 - https://science.mnhn.fr/institution/mnhn/collection/p/item/p06903078 - P 06903078.jpg - hb. E.G. & Aimée Camus - http://www.gbif.org/occurrence/731330495 - https://explore.recolnat.org/occurrence/E5C423F1DBCE487BA640052AD652987A; rev. Ó. Sánchez Pedraja Nov 2016 [the sheet contains five specimens and all correspond to O. hederae]). Bas-Rhin (67): "... / Orobanche Hederae Duby / ... / Naffun, Strasbourg" (https://data.biodiversitydata.nl/naturalis/specimen/L.2817397 - L .2817397.jpg - https://www.gbif.org/occurrence/2516309707; rev. Ó. Sánchez Pedraja Aug 2023). Hauts-de-Seine (92): "11232 / ... Orobanche Hederae Duby / [sur Aralia japonica Thunb (Fatsia japonica (Thunb.) Decne. & Planch. )] / -8 juillet 1973- / jardins Kahn-Boulogne / (Hauts-de-Seine) / -herb. H. Bouby-" (http://coldb.mnhn.fr/catalognumber/mnhn/p/p00598694 - P 00598694.jpg - http://www.gbif.org/occurrence/437165371 - https://explore.recolnat.org/occurrence/9E9554256701499C9B22F50127422C03; rev. Ó. Sánchez Pedraja Feb 2012). Dordogne (24): "Elle vient d'être retrouvée par M. Charles Des Moulins, dans le Périgord" (Vaucher 1827: 57, loc. class. of O. du Lierre); "à Lanquais comme aux environs de Libourne et de Bordeaux" (Des Moulins 1835: 80, loc. class. of O. hederae); Bergerac, Bournazel (pr. Lanquais), 00º 40’ 30,44’’ E, 44º 49’ 03,56’’ N, 65 m, parásita de Hedera sp. en cuneta herbosa sombreada en el borde del camino de entrada sur al castillo de Lanquais, G. Moreno Moral, 9-VI-2012 (obs.). Gironde (33): "aux environs de Libourne. Elle fleurit dans le mois de Juin." (Vaucher 1827: 57, loc. class. of Orobanche du Lierre); "à Lanquais comme aux environs de Libourne et de Bordeaux" (Des Moulins 1835: 80, loc. class. of O. hederae); "Herbier C.n Sipière / Orobanche Hederae Vauch / Libourne / Réc. par Sipière le 12 Juin 1854" (https://science.mnhn.fr/institution/um/collection/mpu/item/mpu462361 - MPU 462361.jpg - http://www.gbif.org/occurrence/1807515810; rev. Ó. Sánchez Pedraja Aug 2023). Vienne (86): "M. Chaboisseau a récolté, sur les racines de l'Angelique des jardins cultivée au séminaire de Montmorillon, une Orobanche que M. Schultz rapporte avec hésitation à une espece de la Grèce qu'il a publiée dans le Prodromus" (Lacroix 1860: 565, loc. class. of O. chaboissaei). Occitanie / Occitania [Languedoc-Roussillon-Midi-Pyrénées] Haute-Garonne (31): "H. Sur les racines du lierre. R. R. Près de Toulouse au Vernet, dans le parc de l'ancien château. C. C. C." (Noulet 1837 : 487, sub O. vaucherii). Hérault
(34): 1.
"Orobanche laurina Godr.! an Bert. / Jardin du plantes 4 juillet 1804
/ Sur le lierre! ..." – 2. "Orobanche
Sarthe (72): "Herbier L. Chevalier / Orobanche procera ... / Environs de Saosnes / dans le médiane ... champ sur / Carduus nutans / 29.6.1876 / ..." (MNHN-P-P04405584 - P 04405584.jpg; rev. Ó. Sánchez Pedraja Mar 2013 [the specimens seem to correspond to O. hederae]). Bouches-du-Rhône
(13):
Aix-en-Provence,
junto al acueducto de Roquefavour [aqueduc
de Roquefavour, Ventabren],
G. Gómez Casares &
G. Moreno Moral, 01.06.2006 (obs.); "Herbier
de M. L'Abbé Lelièvre /
Orobanche
Vaucluse (84): D. 6.6.1999 Ravin de Fouix / Viens (Vs) (https://science.mnhn.fr/institution/mhnaix/collection/aix/item/aix026186 - AIX 026186.jpg; rev. Ó. Sánchez Pedraja Apr 2014 [the specimens seem to correspond to O. hederae]). Georgia / Sakartvelo / Georgien / Géorgie / Грузия (former Georgian Soviet Socialist Republic [Georgian SSR] / Грузинская Советская Социалистическая Республика [Грузинская ССР]) (Beck 1930: 221, sub O. Hederae Duby; Gagnidze 1999; Schneeweiss & al. 2004; Carlón & al. 2005: 64, sub O. hederae Duby; Tarkhnishvili & Chaladze 2013, sub O. hederae Duby; Piwowarczyk & al. 2023: 71-73, sub O. hederae Duby) Adjara / Аджария / Autonomous Republic of Adjara / Аджарская Автономная Республика (Piwowarczyk & al. 2023: 71, sub O. hederae Duby). Imeretia / Имеретия [Imeretiya / Imereti Mchare] (Piwowarczyk & al. 2023: 73, sub O. hederae Duby): Sataplia Managed Reserve [42°18'56.3"N 42°40'24.0"E 42.315626, 42.673319, near Jimastaro], 15 Jun 2021 (https://www.inaturalist.org/observations/83617370 by Michael Darling; accessed Jun 2022 [on Hedera helix aggr.]); Imeretien [42°16'38.7"N 42°42'18.6"E 42.277405, 42.705171, Kutaisi / Кутаиси], 8 Jun 2022 (https://www.inaturalist.org/observations/120794481 by Jacek Pietruszewski in iNaturalist; accessed Jun 2022 [yellow form, on Hedera helix aggr.]. Kakheti / Кахетия (Piwowarczyk & al. 2023: 73, sub O. hederae Duby): "https://www.ncbi.nlm.nih.gov/nuccore/AY209273: Georgia, Great Caucasus, forest reserve of Lagodechi [Lagodekhi Nature Reserve], 600-800 m a.s.l.; on Hedera helix (Araliaceae), GS, AT, MS & PS (WU: S&T 7889)" (https://www.ncbi.nlm.nih.gov/nuccore/29123713) (Schneeweiss & al. 2004; Carlón & al. 2005: 64, sub O. hederae Duby; WU: herb. Schonswetter & Tribsch 7889 https://www.ncbi.nlm.nih.gov/nuccore/83745312 (DQ310019) (Park & al. 2007); "Kachetie, Gruzie [41°51'35.4"N 46°17'28.7"E 41.859839, 46.291314, Lagodekhi Protected Areas / Лагодехи (национальный заповедник)]", 30 Jul 2013 (https://www.inaturalist.org/observations/101043036 - https://www.gbif.org/es/occurrence/3415601432 by Vladimír Fuka in iNaturalist; accessed Jun 2022 [on Hedera helix aggr.]); Tbilisi [41°52'27.3"N 46°14'22.5"E 41.874237, 46.239570, near Khizabavra], 18 Jun 2022 (https://www.inaturalist.org/observations/122341150 - https://www.gbif.org/es/occurrence/3415601432 by Giorgi Natsvlishvili; accessed Jun 2022 [on Hedera helix aggr.]). Racha-Lechkhumi and Kvemo Svaneti / Рача-Лечхуми и Нижняя Сванетия (Piwowarczyk & al. 2023: 73, sub O. hederae Duby). Samtskhe-Javakheti / Самцхе-Джавахети (Piwowarczyk & al. 2023: 73, sub O. hederae Duby). Tbilisi / Тбилиси (Piwowarczyk & al. 2023: 73, sub O. hederae Duby). Abkhazia / Abjasia / Аҧсны / Абхазия / Abkhazie / Abchasien [Aphsny / Apjazeti / Abjazia] (former Abkhaz Autonomous Soviet Socialist Republic [Abkhaz ASSR] / Абхазская Автономная Советская Социалистическая Республика [Абхазская АССР]) (Piwowarczyk & al. 2023: 71, sub O. hederae Duby) "Transkaukasien! in Abchasien (Albow!), im Passe Petznie auf Hedera colchica C. Koch im August 1904 (Woronow!)" (Beck 1930: 221, sub O. Hederae Duby); Пицунда [Pitsunda / Bichvinta / Пиҵунда, 43°09'23.9"N 40°19'45.7"E - 43.156639, 40.329354 (Gagra District / Гагрский район)], 10 Jun 2012 (https://www.inaturalist.org/observations/104574664 - https://www.gbif.org/es/occurrence/3456240074 by Gennadiy Okatov in iNaturalist; accessed Jun 2022 [very illustrative photographs]); Пицунда [Pitsunda / Bichvinta / Пиҵунда, 43°09'23.9"N 40°19'45.7"E - 43.156639, 40.329354 (Gagra District / Гагрский район)], 10 Jun 2012 (https://www.inaturalist.org/observations/104574665 - https://www.gbif.org/es/occurrence/3456251001 by Gennadiy Okatov in iNaturalist; accessed Jun 2022 [on Hedera helix aggr.]). Germany / Deutschland / Alemania / Allemagne [native and naturalized / garden weed] (Beck 1890: 261, sub O. Hederae Duby; Beck 1930: 220, sub O. Hederae Duby; Pusch 2009: 64 Abb. 80, sub O. hederae Duby) Baden-Württemberg / Baden-Württemberg: "Herbarium Dr. Jermstad / Orobanche hederae / Duby / Juli, 1925 / Iskeiner Keotz [Lörrach (district)], Baden" (https://www.unimus.no/felles/bilder/web_hent_bilde.php?id=12971014&type=jpeg - O:V:2254758 - https://www.gbif.org/occurrence/1702126909; rev. Ó. Sánchez Pedraja Aug 2023). Boroughs and neighborhoods of Berlin [ naturalized]: "Berlin [52°27'22.0"N 13°18'24.6"E 52.456099, 13.306824, Gewächshäuser Botanischer Garten Berlin], DE-BE-BR, 5 Jul 2023" (https://www.inaturalist.org/observations/171358592 by Fabian Battiege in iNaturalist; accessed Aug 2023). Lower Saxony / Niedersachsen [naturalized]: "Botanisher Garten [TU Braunschweig Botanical Garden, 52°16'14.9"N 10°31'57.8"E 52.270803, 10.532710], 38106 Braunschweig, 29 Jun 2023" (https://www.inaturalist.org/observations/170134366 - https://www.gbif.org/occurrence/4145462282 by Walter Wimmer in iNaturalist; accessed Aug 2023). Rheinland-Pfalz / Rhineland-Palatinate: "Orobanche hederae Dub / Auf Hedera Helix zu / Oberlahnstein bei Coblenz [Koblenz] / Wirtgen" (http://www.botanicalcollections.be/specimen/BR0000031266284 - BR 0000031266284 - hb. Dieudonne - https://oxalis.br.fgov.be/images/BR0/000/031/266/284/BR0000031266284.jpg - https://www.gbif.org/occurrence/4072832910; rev. Ó. Sánchez Pedraja Aug 2023); idem (https://www.botanicalcollections.be/specimen/BR0000031266277 - BR 0000031266277 - hb. Dieudonne - https://oxalis.br.fgov.be/images/BR0/000/031/266/277/BR0000031266277.jpg - https://www.gbif.org/occurrence/4072398798; rev. Ó. Sánchez Pedraja Aug 2023); "Herbarium Universitatis Mosquensis (MW) / Orobanche hederae Duby / ... / St. Goarshausen / slope of Rhein valley, / xeric forest, glade, on Hedera helix / Соll. A. Seregin, I. Seregina / Nº E-1803 / 12.09.2015" (https://plant.depo.msu.ru/open/public/item/MW0797646 - MW 0797646.jpg; rev. Ó. Sánchez Pedraja Jul 2020). Thüringen / Thuringia: "Germania: ad arcem Liebenstein [Thuringia] prope Kamp ad fl. Rhenum (Spiessen) in exemplaribus pluribus" (Beck 1922: 40[472], loc. class. of O. hederae f. immaniflora). "Die Efeu-Sommerwurz Orobanche hederae Vaucher ex Duby ist ein Vollschmarotzer und kommt als atlantisch-mediterrane Art in West- und Südeuropa vor. In Deutschland erreicht sie am Rhein und in den Tälern von Ahr und Mosel die Ostgrenze ihres natürlichen Areals (Pusch und Günther 2009, Gausmann und Büscher 2011). Ihre Vorkommen im restlichen Deutschland gelten als neophytisch, wobei sie auffällig häufig aus Universitätsstädten gemeldet wurde. Die Art lässt sich leicht auf ihrer weit verbrei-teten Wirtspflanze Efeu ansiedeln und wird deshalb gern in Botanischen Gärten kultiviert, wo sie sich unter günstigen Umständen lange halten und weiter ausbreiten kann. Die in Floraweb und im deutschen Verbreitungsatlas (Netzwerk Phytodiversität 2013) verzeichneten Vorkommen in Kiel, Greifswald, Potsdam, Berlin-Dahlem, Bonn, Münster, Jena und Würzburg etwa dürften sich allesamt auf Botanische Gärten beziehen." (Brandt & Poppendieck 2016: 71, sub O. hederae). Greece / Grecia / Ελλάδα / Hellas (Beck 1890: 261, sub O. Hederae Duby; Beck 1930: 221, sub O. Hederae Duby; Pusch 2009: 64 Abb. 80, sub O. hederae Duby) Continental / Central Greece Στερεά / Κεντρική Ελλάδα Attica Region / Περιφέρεια Αττικής "Graecia: prope Athenas" (Beck 1890: 261, sub O. Hederae Duby). Central Macedonia / Κεντρική Μακεδονία Kilkis (regional unit) / Περιφερειακή ενότητα Κιλκίς: "Axioupoli [Axioupoli / Αξιούπολη / Boymitsa / Боймица / Μποέμιτσαi, 41°03'43.6"N 22°23'35.2"E 41.062114, 22.393108, Gripas (Fanou), Peonia / Παιονία, 8 Jul 2023" (https://www.inaturalist.org/observations/173647094 by Leon Orcot in iNaturalist; accessed Jul 2023). "Rare on Crete, known from only a few locations" (O. hederae in Cretan Flora, accessed Jul 2020). Heraklion / Περιφερειακή ενότητα Ηρακλείου: "Nikos Kazantzakis [35°14'28.2"N 25°13'15.4"E 35.241153, 25.220938, Astrakiano Gorge (Archanes-Asterousia)], 18 May 2023" (https://www.inaturalist.org/observations/162364329 - https://www.gbif.org/es/occurrence/4120992171 by https://www.inaturalist.org/people/mcusson in iNaturalist; accessed Aug 2023). North Aegean Region / Περιφέρεια Βορείου Αιγαίου Chios / Χίος / Khíos / Híos (https://www.gbif.org/es/occurrence/788717613 [n.v.]). Samos / Σάμος: "... Collected by: A.J. Winter / no. 14 d.d.: 25 May 1995 / Greece, Sámos, ½ km S of Kokkari / 37°47"N, 26°53"E Alt: 150 m / ... on Hedera..." (https://data.biodiversitydata.nl/naturalis/specimen/WAG.1141305 - WAG.1141305.jpg; rev. Ó. Sánchez Pedraja Jul 2020); "Samos, Karlovasi [37°46'22.2"N 26°39'54.0"E 37.772825, 26.664990], 24 May 2010" (https://www.inaturalist.org/observations/68248889 - https://www.gbif.org/es/occurrence/3031904133 by David Lowry in iNaturalist; accessed Jul 2023). Hungary / Magyarország / Hungría / Hongrie / Ungarn [naturalized, needs be confirmed] (Uhlich & al. 1995: 116, sub O. hederae; Pusch 2009: 64 Abb. 80, sub O. hederae Duby) Veszprém County: "Ungarn: selten, Csopak, Somlo-hegy [Somló,Veszprém County], Badacsonyi [Badacsony, Badacsonytomaj] (leg. Uhlich 1981)" (Uhlich & al. 1995: 116, sub O. hederae). Ireland / Éire / Irlanda / Irlande (Beck 1930: 220, sub O. Hederae Duby; Rumsey & Jury 1991: 272-273, sub O. hederae Duby; Pusch 2009: 64 Abb. 80, sub O. hederae Duby; map by BSBI, accessed Jul 2020; Zoë Devlin 2008-2020 in Wild flowers of Ireland, accessed Jul 2020) Donegal County: "Whitehill [54°48'27.6"N 7°41'21.8"W 54.807662, -7.689387], The Bower, 2 Ene 2005" (https://www.inaturalist.org/observations/34450015 by Rhonda Ridley in iNaturalist; accessed Aug 2023). Kerry County: "in the Killarney area of Ireland" (Rumsey & Jury 1991: 272, sub O. hederae Duby). Italy / Italia / Italien (Vaucher 1827: 57, loc. class. of O. du Lierre; Bertoloni 1846: 423, loc. class. of O. yuccae; Bertoloni 1846: 424, loc. class. of O. laurina; Reuter 1847: 28, sub O. Hederae Duby; Reuter 1847: 719, loc. class. of O. glaberrima; Facchini 1855: 76, loc. class. of O. cerina; Beck 1890: 260, loc. class. of O. Helicis Rota; Beck 1890: 261, O. Hederae Duby; Beck 1930: 221, sub O. Hederae Duby; Pusch 2009: 64 Abb. 80, sub O. hederae Duby) Naples
(NA): 1.
"Orobanche glaberrima
... [description] ...".
– 2. "Orobanche glaberrima Guss.!
/ Reut. in DC. prod. / [m. Reuter]".
– 3. "O. glaberrima
[m. Reuter]
/
Scripsit Metropolitan City of Rome Capital: "Ann. Habui Româ [Rome] ab Ill. Principe de Canino, et Musiniano, qui detexit parasiticam supra radices Lauri nobilis L." (Bertoloni 1846: 424, loc. class. of O. laurina Bonap. ex Bertol.) Bergamo (BG): "Orobanche Helicis Rota, Flora della provincia di Bergamo [Bergamo, Italy] p. 101 ... Vidi sepc. origin. in herb. Cesati." (Beck 1890: 260, loc. class. of O. Helicis Rota; Beck 1930: 219), "Bergamo" (Beck 1930: 221, sub O. Hederae Duby). Savona (SV): "Cette singulière Orobanche se trouve sur les collines qui s'étendent de Gênes à Savone [Genoa / Genova to Savone / Savona]. Je l'ai cueillie en 1825, sur les hauteurs qui dominent cette dernière ville, et dans les maisons de campagne du Ponente. Je l'ai revue l'année suivante en grande abondance à Gênes même, dans les jardins Négri" (Vaucher 1827: 57, loc. class. of O. du Lierre). Genoa (GE): "Italien! verbreitet bei Genua [Genoa / Genova]" (Beck 1930: 221, sub O. Hederae Duby); Rapallo, No. 1099 A. Schoenmakers 30.5.1932 (https://www.gbif.org/es/occurrence/2517043542 - U.1469250.jpg; rev. Ó. Sánchez Pedraja Jul 2020). Trentino-Alto Adige / Südtirol South Tyrol / Bolzano (BZ): "Orobanche cerina / Koch ined. / Ober Bench nordlichsim der Nähe von Botzen [?] / Dr. Facchini" (https://data.biodiversitydata.nl/naturalis/specimen/L.2817685 - L 2817685.jpg, rev. Ó. Sánchez Pedraja Nov 2020); "Postea satis frequenter crescentem plantam islam ad Hederae Helicis radices in ditione Bulsanensi [Bolzano / Bozen] inveni, quam cum O. minore conjungo" (Facchini 1855: 76, loc. class. of O. cerina). Pisa (PI): "Ann. Habui ex sylva Pisana [Pisa] prope mare, et ex horto botanico Pisano, quibus in locis reperta est parasitica supra radices Yucccae aloifoliae L., a Prof. Petro Savio" (Bertoloni 1846: 423, loc. class. of O. yuccae); "Flora Etrusca Exsiccata / Orobanche minor Sutton / Agro pisano, ... radicii dell' Hedera Helix / Fior Maggio / 1847 / ex. Herb. Hor. bot. Pisani" (MNHN-P-P04392695 - P 04392695.jpg; rev. Ó. Sánchez Pedraja Jan 2014); "Flora Etrusa Exsiccata / Orobanche minor Sutton / Bocca d'Arno [Boccadarno], Tomboli / Sulle Leguminose [?] / Maggio 25 / 1862 ex Herb. Hort. bot. Pisani" (https://science.mnhn.fr/institution/mnhn/collection/p/item/p04384944 - P 04384944.jpg - http://www.gbif.org/occurrence/464879266 - https://explore.recolnat.org/occurrence/C3853D0D023946FBA4878D33D967E340; rev. Ó. Sánchez Pedraja Feb 2012 [corolla-tube constricted towards its distal end]). Sardinia / Sardegna / Cerdeña (photos in www.Sardegnaflora.it, accessed Jul 2020) "Sardinien! (Arizzo, Laconi, Sarcidano)" (Beck 1930: 221, sub O. Hederae Duby). Palermo (PA): "Sizilien! (Ficuzza, Palermo)" (Beck 1930: 221, sub O. Hederae Duby). Liechtenstein (Beck 1930: 221, sub O. Hederae Duby; Pusch 2009: 64 Abb. 80, sub O. hederae Duby) Vaduz: "Liechtenstein! bei Vaduz" (Beck 1930: 221, sub O. Hederae Duby). Luxembourg / Lëtzebuerg / Luxembourg / Luxemburg / Luxemburgo (Uhlich & al. 1995: 116, sub O. hederae; Colling 2005: 40, "R - N - FOR", sub O. hederae Duby; Pusch 2009: 64 Abb. 80, sub O. hederae Duby) Echternach County: "1987 selten bei Echternach, on noch ?" (Uhlich & al. 1995: 116, sub O. hederae). Image Code: ORBHE-0. ORBHE-1. ORBHE-2. ORBHE-3. ORBHE-4. ORBHE-5. (Mifsud, sub O. hederae in Malta Wild Plants, accessed Apr 2013). Montenegro / Crna Gora [Black Mountain] / Црна Гора / Черногория (Pulević 2005) Bar Municipality / Opština Bar / Општина Бар: "Добра-Вода [Dobra Voda], Черногория, 21 May 2023" (https://www.gbif.org/es/occurrence/4121394591 - https://www.inaturalist.org/observations/162979969 by Roman Romanov in iNaturalist; accessed Aug 2023); "Dupilo [42°15'17.7"N 19°02'19.4"E 42.254921, 19.038727], 27 May 2023" (https://www.gbif.org/occurrence/4127106515 by https://www.inaturalist.org/people/donnmaier in iNaturalist; accessed Aug 2023). Kotor Municipality Opština / Kotor Општина Котор: "9Q56+FH9, Радановићи [Radanovići, 42°21'29.2"N 18°45'42.4"E 42.358114, 18.761767], Черногория, 20 May 2023" (https://www.inaturalist.org/observations/162609607 by Anfisa Yurenskaya in iNaturalist; accessed Aug 2023). Netherlands / Nederlanden / Países Bajos / Pays-Bas (Pusch 2009: 64 Abb. 80, sub O. hederae Duby) Friesland Province / Frisia: "Passage [53°13'04.8"N 5°47'22.1"E 53.217995, 5.789480], Leeuwarden, 16 Jul 2020" (https://www.inaturalist.org/observations/53403938 by Bram Scholma in iNaturalist; accessed Aug 2023). North Macedonia / Северна Македонија / Macedonia del Norte / Macédoine du Nord / Nordmazedonien (Pusch 2009: 64 Abb. 80, sub O. hederae Duby; Nikolov 2020: 22, sub O. hederae Duby) Skopje Statistical Region / Скопски Регион: "1. Matka (Skopje [Скопје, Municipality of Sopište / Општина Сопиште]): in the vicinity of the dam «Matka», along the narrow path that leads to the caves, 343 m a.s.l., 10.06.2011 (N: 42º 00’ 37.8’’; E: 021º 30’ 46.9’’) (Leg./Det.: Z. Nikolov)" (Nikolov 2020: 22, sub O. hederae Duby). Southeastern Statistical Region / Југоисточен Регион: "Mt. Belasica [Беласица]: village Banjsko [Банско] (Rudski, 1943)" (Nikolov 2020: 22, sub O. hederae Duby). Southwestern Statistical Region / Југозападен Регион: "2. Debar (village Modrič [Модрич]): in the valley of the river Crn Drim, 594 m.a.s.l., 22.06.2012 (Leg./Det.: Z. Nikolov)" (Nikolov 2020: 22, sub O. hederae Duby). Norway / Norge / Noreg / Noruega [naturalized] (Lid & Lid 1994; Pusch 2009: 64 Abb. 80, "synanthrop", sub O. hederae Duby; Gederaas & 2012: 123, sub O. hederae) Eastern Norway / Østlandet / Austlandet: "Østfold" (Gederaas & 2012: 123, "NK", sub O. hederae). Poland / Polska / Polonia / Polen [naturalized] (Uhlich & al. 1995: 116, sub O. hederae) Opole Voivodeship: "in Schlesien [Silesia] in Proskov [Prószków / Proskau] bei Opole eingeschleppt, on noch ?" (Uhlich & al. 1995: 116, sub O. hederae); "Bardzo rzadko zawlekana w parkach (Gryfino [West Pomeranian], Pruszków k. Opola). Pasożytuje na bluszczu pospolitym (Hedera helix)" (https://atlas-roslin.pl/gatunki/Orobanche.htm#KLUCZE-5573 - https://atlas-roslin.pl/gatunki/Orobanche_hederae.htm in Flora Polski·atlas-roslin.pl; accessed Aug 2023). Portugal (Willkomm 1870: 625, sub O. Hederae Duby; Colmeiro 1888: 279, sub O. Hederae Dub.; Beck 1890: 261, sub O. Hederae Duby; Guimarães 1904: 179, sub O. Hederae Duby; Beck 1930: 220, sub O. Hederae Duby; Foley 2001: 61, sub O. hederae Vaucher ex Duby; Pusch 2009: 64 Abb. 80, "Fundort ungenau", sub O. hederae Duby) Alto Alentejo (AAl): "AAl" (Foley 2001: 61, sub O. hederae Vaucher ex Duby). Algarve (Ag): "(Ag)" (Foley 2001: 61, sub O. hederae Vaucher ex Duby); "8150, Portugal [37°10'43.4"N 7°56'04.2"W 37.178711, -7.934508, São Brás de Alportel (Faro)], 12 May 2021" (https://www.inaturalist.org/observations/79847347 by Joao Tiago Tavares; accessed et rev. Ó. Sánchez Pedraja Aug 2023, O. minor, saltem p.p. [purple stigma]). Azores / Açores [see Macaronesia / Macaronésie / Makaronesien] Baixo Alentejo (BA): "(BA)" (Foley 2001: 61, sub O. hederae Vaucher ex Duby). Beira Alta (BAl): "(BAl)" (Foley 2001: 61, sub O. hederae Vaucher ex Duby). Beira Baixa (BB): "(BB)" (Foley 2001: 61, sub O. hederae Vaucher ex Duby). Beira Litoral (BL): "Espinhal [Coimbra District] (A.J. Freiré!). Centro littoral:—Arredores de Coimbra (A. de Carvalho!), Santa Cruz (Lopes Manita!), cerca do Jardim Botánico (Loureiro!), muro da Penitenciaria (M. de Lucena!)" (Guimarães 1904: 181, sub O. hederae var. trichodea); "(BL)" (Foley 2001: 61, sub O. hederae Vaucher ex Duby). Douro Litoral (DL): "(DL)" (Foley 2001: 61, sub O. hederae Vaucher ex Duby). Estremadura (E): "Lisboa (Pimentel)" (Colmeiro 1888: 279, sub O. Hederae Dub.); Sintra (Beck 1930: 220, sub O. Hederae Duby); "E" (Foley 2001: 61, sub O. hederae Vaucher ex Duby). Minho (Mi): "Alemdouro littoral:—Valença: [Viana do Castelo District] Costa da E'rvilha (R. da Cunha!)" (Guimarães 1904: 181, sub O. hederae var. trichodea); "(Mi)" (Foley 2001: 61, sub O. hederae Vaucher ex Duby). Ribatejo (R): "(R)" (Foley 2001: 61, sub O. hederae Vaucher ex Duby). Trás-os-Montes (Alto Douro) (TM): "Braganza [Bragança] (Oliveira)" (Colmeiro 1888: 279); "(TM)" (Foley 2001: 61, sub O. hederae Vaucher ex Duby). Romania / România / Rumanía / Rumänien Bucharest: "Bucureşti Sectorul 6, Bucharest [44°26'14.7"N 26°03'42.1"E 44.437418, 26.061681], 9 Jun 2021" (https://www.inaturalist.org/observations/82265099 by Radu Popescu Vega in iNaturalist; accessed Jun 2022). Russia / Россия / Rusia / Russland / Russie (Novopokrovskij & Tzvelev 1958: 98[104], sub O. hederae Duby; Pusch 2009: 64 Abb. 80, "Fundort ungenau", sub O. hederae Duby) Central Federal District / Центральный федеральный округ [unconfirmed] Voronezh Oblast / Воронежская область [unconfirmed]: "Новохоперский р-н [Novokhopyorsky District, 51°17'31.8"N 41°30'01.0"E 51.292164, 41.500267], 6 Ago 2022" (https://www.inaturalist.org/observations/129666553 by Valentina Kostina in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Jul 2023 [O. cernua aggr. (O. cf. cumana)]). North Caucasian Federal District / Северо-Кавказский федеральный округ Dagestan / Daghestan / Республика Дагестан: "Приморское [Samur / Самур, 41°51'16.5"N 48°30'00.1"E 41.854578, 48.500017], Дагестан, РФ - 13 Jun 2020" (https://www.inaturalist.org/observations/49754370 - https://www.gbif.org/es/occurrence/2803071115 by Ramazan Murtazaliev in iNaturalist; accessed Jul 2020); "Республика Дагестан [Gosudarstvennyy Prirodnyy Zakaznik Federal'nogo Znacheniya "Samurskiy" (Государственный природный заказник федерального значения "Самурский"), 41°51'35.8"N 48°33'07.4"E 41.859934, 48.552065]" (https://www.inaturalist.org/observations/27680871 - https://www.gbif.org/es/occurrence/2273303267 by Elena Ilyna in iNaturalist; accessed Jul 2020). Northwestern Federal District / Северо-Западный федеральный округ Kaliningrad Oblast / Калининградская область [alien?]: "Ботанический сад БФУ им. И. Канта [Botanicheskiy Sad Baltiyskogo Federal'nogo Universiteta Imeni Immanuila Kanta, 54°44'13.1"N 20°30'59.5"E 54.736978, 20.516537, Kaliningrad / Калининград / Königsberg], 21 Jul 2022" (https://www.inaturalist.org/observations/127215583 by Юлия Гришанова / Julia Grishanova in iNaturalist; accessed Aug 2023). Southern Federal District / Южный федеральный округ (Pusch 2009: 64 Abb. 80, "Fundort ungenau", sub O. hederae Duby) Krasnodar Krai / Краснодарский край / Krasnoyarsk Krai / Krasnoyarsky kray / Krasnodarsky kray:: "... / Orobanche cumana Wallr. / Краснодарский край, Адлерский район [Krasnodar Territory, Adlersky City District / Адлерский внутригородской район] / левый берег р. Мзымта [Mzymta] ниже пос. Монастырь / (Красная Скала) [(Krasnyy Gostagay), Anapsky District / Анапский район] / буковый самшитовый лес ниже устья Глубокого [mouth of Glubokoe] ... / Собр. А. Серёгин, Л. Ашуркова, З. Винокурова [Coll. A. Seregin et al.] / 12.09.2008. N.º C-1079" (https://plant.depo.msu.ru/open/public/item/MW0723461 - MW 0723461.jpg; rev. Ó. Sánchez Pedraja Jul 2018 [the specimens seem to correspond to O. hederae]); г. Новороссийск [Novorossiysk, 44°43'46.6"N 37°26'41.4"E 44.729599, 37.444820, near Malyi Utrish / Малый Утриш], 28 Jan 2009 (https://www.inaturalist.org/observations/19712901 - https://www.gbif.org/occurrence/2005276449 by Maxim Nuraliev in iNaturalist; accessed Jun 2022); "г.Новороссийск [Novorossiysk, 44°43'44.7"N 37°26'39.0"E] - 2 Jun 2013" (https://www.inaturalist.org/observations/19728535 by Maxim Nuraliev in iNaturalist; accessed Jul 2020); "... Orobanche caryophyllacea Smith / Краснодарский край, Анапский район [Anapsky District], / ... Абрау [Abrau], Навагирский хребет [Navagirskiy Khrebet], Лобанова щель [Lobanova shchel'] ... 650 m ... / 44°42'42.7"N 37°28'44.9"E / на Carpinus orientalis [erroneous host-plant]. / 26.05.2018 / Рябенко О.И. [Ryabenko O.I.] ..." (https://plant.depo.msu.ru/open/public/item/MW1008919 - MW 1008919.jpg; rev. Ó. Sánchez Pedraja Mar 2021 [O. hederae]); "Цветущие растения. Краснодарский край, м/о город-курорт Сочи [Sochi], окр. с. Хлебороб [Khleborob, 43°33'24.5"N 39°52'44.1"E 43.556806, 39.878903], лиственный лес. 12 июня 2017 г." (https://www.plantarium.ru/page/image/id/516141.html, https://www.plantarium.ru/page/image/id/516142.html, https://www.plantarium.ru/page/image/id/516143.html, https://www.plantarium.ru/page/image/id/516144.html, https://www.plantarium.ru/page/image/id/516145.html and https://www.plantarium.ru/page/image/id/516146.html by Александр Фатерыга [Alexander Fateryga] in Plantarium; accessed Aug 2023). Krim / Crimea / Крым / Крим / Tauric Peninsula [see Ukraine / Russia] (Beck 1890: 261, sub O. Hederae Duby; Novopokrovskij & Tzvelev 1958: 98[104], sub O. hederae Duby) Serbia / Srbija / Србијa / Serbie / Serbien [needs be confirmed] (Josifović 1970-1977; Kojić & al. 2001, sub O. hederae Duby; Pusch 2009: 64 Abb. 80, sub O. hederae Duby) Slovakia / Slovensko / Eslovaquia / Slowakei [unconfirmed] Slovenia / Slovenija / Eslovenia / Slowenien (Beck 1890: 261, sub O. Hederae Duby; Martinčič & al. 1999; Pusch 2009: 64 Abb. 80, sub O. hederae Duby) Coastal–Karst Statistical Region / Obalno-kraška statistična regija Slovene Istria / slovenska Istra / Istria slovena: "Capo d'Istria [Koper / Capodistria / Kopar]" (Beck 1890: 261, sub O. Hederae Duby). Slovene Littoral / Primorska / Litorale: "[45°44'19.4"N 13°49'08.1"E 45.738724, 13.818912, Dutovlje / Duttogliano], Jun 2023" (https://www.inaturalist.org/observations/165753921 by brandy midura in iNaturalist; accessed Jul 2023). Spain / España / Espagne / Spanien / Испания (Duby 1828: 350, loc. class. of O. hederae; Reuter 1847: 28, sub O. Hederae Duby; Willkomm 1870: 625, sub O. Hederae Duby; Colmeiro 1888: 279, sub O. Hederae Dub.; Beck 1890: 261, sub O. Hederae Duby; Beck 1930: 220, sub O. Hederae Duby; Aseginolaza & al. 1985: 704; Foley 2001: 61, sub O. hederae Vaucher ex Duby; Pusch 2009: 64 Abb. 80, sub O. hederae Duby) Álava / Araba (Vi): Arceniega, 30TVN8974, 305 m, 22/06/1980, P.M. Uribe-Echebarría (VIT 33076, sub O. hederae [rev. para «Contribuciones VI» / O. hederae Vaucher ex Duby / rev. Ó. Sánchez Pedraja, 4-IX-2002]) (Carlón & al. 2002: 27, sub O. hederae); "«Amarita [Amárita]», 17-VII-1978, (herb. Alejandre 445/78, sub O. gr. minor; rev. Ó. Sánchez Pedraja, 4-IX-2002); ibidem, 4-VIII-1980 (herb. Alejandre 2296/80, sub O. gr. minor; rev. Ó. Sánchez Pedraja, 4-IX-2002); «Maestu / Corres, Manchibrio», 30TWN4528, 900-935 m, P.M. Uribe-Echebarría, 11-VII-2000 (VIT 63584!) y citas (Aseginolaza & al. 1985: 704), éstas parcialmente confirmadas por VIT (33076, «Arceniaga», y 33079, «Zurbano»!)" (Carlón & al. 2002: 26, sub O. hederae); Hueto Abajo, 30TWN1548, 560 m (http://www.herbario.ian-ani.org/index.php?c=pliegos&a=ficha&i=1366 by Pello Urrutia in herbario digital Xavier de Arizaga, Instituto Alavés de la Naturaleza, accessed Jan 2012). Albacete (Ab): Fuensanta, del Carrasco a Fuensanta 3,6 km, 30SWJ8242, 650 m (Molina Cantos & al. 2008). Alicante / Alacant (A): "A" (Foley 2001: 61, sub O. hederae Vaucher ex Duby); "jardines de Alcoi (Cámara 1942: 336) ... llegando a la Serra del Cid, ya en el subsector alcoyano-villenense (Juan 1995: 115) ... fageca, Serra de Serrella, Plà de la Casa, YH3689, 1350 m, L. Serra, J. Pérez & J.J. Izquierdo, 5-9-2002, v.v. ..." (Serra Laliga 2007: 788); BDBC (accessed Jul 2020); "Alicante" (O. hederae Duby in Herbari Virtual del Mediterrani Occidental, accessed Jan 2012). Almería (Al): "Río Paterna" (photos by Andrés Ivorra in Ivorra & al., Plantas parásitas de Almería; accessed Jun 2013). Asturias [formerly known as Ovideo Province] (O): "Astur. (pr. Gijón, Dur. [Durieu])" (Willkomm 1870: 625, sub O. Hederae Duby; Colmeiro 1888: 279, sub O. Hederae Dub.); "cerca de Oviedo (L.P. Ming.)" (Colmeiro 1888: 279, sub O. Hederae Dub.); "O" (Foley 2001: 61, sub O. hederae Vaucher ex Duby). Barcelona (B): "Bacinonem (Benth.!)" (Reuter 1847: 28, sub O. Hederae Duby); "Catal. (pr. Bacinon., Benth., versus Hortá [Horta, Horta-Guinardó, Barcelona district] ... San Miguel del Fay [monasterio de Sant Miquel del Fai, Riells, Vallés Oriental], Texid." (Willkomm 1870: 625, sub O. Hederae Duby; Colmeiro 1888: 279, sub O. Hederae Dub.); "Monserrat [Montserrat]" (Colmeiro 1888: 279, sub O. Hederae Dub.); "B" (Foley 2001: 61, sub O. hederae Vaucher ex Duby); BDBC ( accessed Jul 2020); "Barcelona" (O. hederae Duby in Herbari Virtual del Mediterrani Occidental, accessed Jan 2012). Baleares, Islas / Balears, Illes / Balearic Islands [formerly known as Palma de Mallorca Province ("PM")] (Foley 2001: 61, sub O. hederae Vaucher ex Duby, "PM"; BDBC ( accessed Jul 2020). Ibiza / Eivissa: "Ibiza" (O. hederae Duby in Herbari Virtual del Mediterrani Occidental, accessed Jan 2012). Majorca / Mallorca: "Baleares en Mallorca (Burnat, Barbey)" (Colmeiro 1888: 279, sub O. Hederae Dub.); "Plantes d'Espagne — F. Sennen / N.º 2018 / Orobanche balearica / Baléares: Alfabia, près de Palma, / sur le lierre / 1914-18-VI Leg. F. Bianor" (https://www.gbif.org/es/occurrence/1840543035 - http:/www.botanicalcollections.be/specimen/BR0000026151717V - BR0000026151717V.jpg et https://data.biodiversitydata.nl/naturalis/specimen/L.2817406 - L.2817406.jpg, rev. Ó. Sánchez Pedraja Jul 2020, syntypes of O. balearica); idem (US 2128316 - Image No. Barcode 03926915 - http://n2t.net/ark:/65665/m34446de17-3a51-4ff3-a054-0ae9d91a2950 - https://www.gbif.org/es/occurrence/2625843870, rev. Ó. Sánchez Pedraja Jun 2022, syntypes of O. balearica); "N.º 2018 / Orobanche balearica Sen et Pau / [ Baléares: Alfabia, près de Palma, sur le lierre] 1914-18-VI / Leg. F. Bianor" (LY 0475996 - hb. R. Bonaparte - https://mediaphoto.mnhn.fr/media/15451552944546CuT76I8Atb1qOmm - https://explore.recolnat.org/occurrence/2FF9139E841B4110BAD5CD0BBC9B5775, rev. Ó. Sánchez Pedraja Jun 2022, syntypes of O. balearica); "Mis amigos, los Hermanos Sennen y Elías, han tenido la bondad de comunicarme unas cuantas plantas, recogidas en las cercanías de Sóller" (Pau 1911: 18, loc. class. of O. balearica); "Mll" (Foley 2001: 61, sub O. hederae Vaucher ex Duby); "Mallorca" (O. hederae Duby in Herbari Virtual del Mediterrani Occidental, accessed Jan 2012). Menorca / Minorca: "(Mn)" (Foley 2001: 61, sub O. hederae Vaucher ex Duby); "Menorca" (O. hederae Duby in Herbari Virtual del Mediterrani Occidental, accessed Jan 2012). Burgos (Bu): "Plantas Españolas / Orobanche hederae Vch. f. exaltata / Castilla: Bujedo [Bugedo]: champs boisés / 17.6 09 [1909] leg. Elias" (https://data.biodiversitydata.nl/naturalis/specimen/U.1469256 - U.1469256.jpg, rev. Ó. Sánchez Pedraja Jul 2020); "Plantas Españolas / Orobanche hederae Vch. f. rosea / Castilla: Bujedo [Bugedo]: sur / le Lierre / 27.6 09 [1909] leg. Hno. Elias" (https://data.biodiversitydata.nl/naturalis/specimen/U.1469257 - U.1469257.jpg, rev. Ó. Sánchez Pedraja Jul 2020); "Plantes d'Espagne / Orobanche hederae Vauch. / Castilla: Bujedo [Bugedo] sur le Lierre / 1909 20.6 Hno. H. Elias" (https://data.biodiversitydata.nl/naturalis/specimen/L.2817409 - L.2817409.jpg, rev. Ó. Sánchez Pedraja Jul 2020); "herbario Laínz de Oña, 30TVN6534, López-Villaseñor, 25-VI-1964" (Carlón & al. 2002: 26, sub O. hederae); bajo Peña Carriá, sobre Arroyo de San Zadornil, 30TVN8545, 1000 m, aparentemente sobre la Hedera helix L. que tapiza el cantil (al pie de cantil calizo), G. Moreno Moral MM0131/2003, 19.06.2003 (herb. Sánchez Pedraja 11438); "VN34: Valle de Sedano..." (Alejandre & al. 2006: 464). Cáceres (Cc): Cabezuela del Valle, 19-VI-1990, leg. J.L. Pérez Chiscano (SALA 86189); Hervás, 30TTK5762, 19-VI-1990, leg. J.L. Pérez Chiscano (SALA 86191); "Cc" (Foley 2001: 61, sub O. hederae Vaucher ex Duby). Cádiz (Ca): "Ca" (Foley 2001: 61, sub O. hederae Vaucher ex Duby). Canarias, Islas / Canary Islands / Kanarische Inseln (Gilli 1980: 115; Reyes, pers. comm., Jul 2020) [see Macaronesia / Macaronésie / Makaronesien] Cantabria [formerly known as Santander Province] (S): "Cantabr. (c. Santander ... Lge. ...)" (Willkomm 1870: 625, sub O. Hederae Duby; Colmeiro 1888: 279, sub O. Hederae Dub.); "Ogarrio (S. Gut.)" (Colmeiro 1888: 279, sub O. Hederae Dub.); "(S)" (Foley 2001: 61, sub O. hederae Vaucher ex Duby); BIO 9637 / Orobanche minor / Monte Buciero, Santoña, 40 m / terreno removido y seco con nitrofilia / 1-5-1987 / M. Herrera // Revisión para «Orobanchaceae - Paleártico» / O. hederae Vaucher ex Duby / Det.-rev. Ó. Sánchez Pedraja, 22.09.2006 [obs. ejemplar atípico. flores aparentemente estrechadas en la garganta; anteras c. 1 mm, grisáceas con apículo blanco]; Medio Cudeyo, Hermosa, 30TVP4002, 40 m, sobre Hedera helix L. [aggr.] (Hederaceae) en una pared, Ó. Sánchez Pedraja, 13.06.1993 (herb. Sánchez Pedraja 02528); Tresviso, entre la central y el primer puente sobre el río Urdón (pr. Urdón), 30TUN6791, 90 m, sobre (?) Hedera helix L. [aggr.] (Hederaceae), G. Gómez Casares, 07.10.2002 (herb. Sánchez Pedraja 10962); Campoo de Yuso, las Carreras (pr. Bustamante), 30TVN1761, 860 m, junto a robles con H. helix L. [aggr.], sotobosque de caducifolios (robledal con hayas, avellanos y acebos), G. Moreno Moral, 23.07.2006 (obs.); Municipality of Santander, Santander City, 30TVP3513, 10 m, parasitic on Hedera sp. [cv. unknown] (Araliaceae) cultivated in the flowerbed, Ó. Sánchez Pedraja & M. Tapia Bon SP0001/2010, 09.05.2010 (herb. Sánchez Pedraja 13489); Cillorigo de Liébana, junto a Viñón, 30TUN6682, 560 m, parásita de H. helix L. [aggr.] en borde de camino, G. Moreno Moral, 26-V-2011 (obs. [a good population which is specially composed for specimens of the yellow form]). Castellón / Castelló (Cs): "Cs" (Foley 2001: 61, sub O. hederae Vaucher ex Duby; BDBC, accessed Jul 2020). Córdoba / Cordova (Co): "(Co)" (Foley 2001: 61, sub O. hederae Vaucher ex Duby). Coruña, La / A Coruña (C): "Cedeira en Silva Pando & al. (2000: 27)!" (Carlón & al. 2002: 26, sub O. hederae). Gerona / Girona (Ge / Gi): "c. San Juan las Fonts [Sant Joan les Fonts, Garrotxa]" (Willkomm 1870: 625, sub O. Hederae Duby); "Talaixá [Caserio Talaixa, Caserío Talaixá] (Vayr.)" (Colmeiro 1888: 279, sub O. Hederae Dub.); Nº 9010 Sant Pere de Rodes 10 Aug 1965 H.C.D. Wit (https://data.biodiversitydata.nl/naturalis/specimen/WAG.1141309 - WAG.1141309.jpg, rev. Ó. Sánchez Pedraja Jul 2020); "Ge" (Foley 2001: 61, sub O. hederae Vaucher ex Duby); Palafrugell, cap de Sant Sebastià (sobre Llafranc, Baix Empordà [Bajo Ampurdán]), 31TEG1638, 150 m, 25.08.2006, sobre (!) Hedera helix L. [aggr.] (Hederaceae) en un pinar, Ó. Sánchez Pedraja & M. Tapia Bon (obs.); BDBC (accessed Jul 2020); "Gerona" (O. hederae Duby in Herbari Virtual del Mediterrani Occidental, accessed Jan 2012); Выкопанные цветущие растения, паразитирующие на Hedera helix (побег слева). Испания, автономное сообщество Каталония, провинция Жирона [Girona], комарка Баш Эмпорда [Baix Empordà], муниципалитет Калонже [Calonge], переходная зона от поймы малой реки к дубовому редколесью. 14.05.2020 (https://www.plantarium.ru/page/image/id/656396.html, id/656398.html, id/656400, id/656402.html and https://www.plantarium.ru/page/image/id/656406.html by Светлана Коурова [Svetlana Kourova] 2020 in Plantarium; accessed Jun 2020). Granada (Gr): "regno Granatensi (Boiss. [Boissier]!)" (Reuter 1847: 28, sub O. Hederae Duby); "regno Granat. (Bss. [Boissier])" (Willkomm 1870: 625, sub O. Hederae Duby; Colmeiro 1888: 279, sub O. Hederae Dub.); "Gr" (Foley 2001: 61, sub O. hederae Vaucher ex Duby). Guadalajara (Gu): "Gu" (Foley 2001: 61, sub O. hederae Vaucher ex Duby). Guipúzcoa / Gipuzkoa (SS): 1. "Willkommii it. hisp. secund. / 183. / Orobanche hederae Vauch. / In radicibus Hederae Helicis prope Villafranca / in Guipuzcoa raro / Majo 1850." (COI 00056890 - http://coicatalogue.uc.pt/specimen/56889, rev. Ó. Sánchez Pedraja Jul 2020); "Villafranca [Villafranca de Ordicia / Ordizia, Goyerri / Goierri] in Guipuzcoa raro Willkomm! It. secund. 183.)" (Reichenbach fil. 1862: 103, sub O. Hederae Duby); "Cantabr. (...Villafranca [Villafranca de Ordicia / Ordizia, Goyerri / Goierri] in Guipuzcoa, Wk.)" (Willkomm 1870: 625, sub O. Hederae Duby; Colmeiro 1888: 279, sub O. Hederae Dub.); "SS" (Foley 2001: 61, sub O. hederae Vaucher ex Duby). Huesca (Hu): "Hu" (Foley 2001: 61, sub O. hederae Vaucher ex Duby). Jaén (J): "(J)" (Foley 2001: 61, sub O. hederae Vaucher ex Duby); Santo Tomé, Collado de Peña Rubia (siguiendo la senda y a poco de trasponer el collado e iniciar la bajada hacia el Cortijo del Calerón, ya en el término municipal de Santo Tomé), 30SWH0903, 900 m, G. Moreno Moral, 9-VI-2013 (obs.). León / Llión (Le): Montes Aquilianos, Ponferrada, San Pedro de Montes, 29TPH9902, 1000 m, sobre Hedera helix, ruinas, 20-VII-1997, s.c. (Hergueta & al. 1999). Lérida / Lleida (L / Ll): Lés (Coste 1922); BDBC (accessed Jul 2020); "Flora Hispanica ... / Orobanche hederae Duby / Prov. Lèrida (Spaanse Pyreneeen); Z. van / Las Bordas (Z.Z.O. van Luchon), bij / Refugio. alt. 1050 m / Leg. R. Horreüs de Haas no. 30 / det K.U. Kramer" (https://data.biodiversitydata.nl/naturalis/specimen/U.1469193 - U.1469193.jpg, rev. Ó. Sánchez Pedraja Jul 2020 [sub O. gracilis]). Lugo (Lu): "Lu" (Foley 2001: 61, sub O. hederae Vaucher ex Duby). Madrid (M): 1. "H. M. Willkommii iter hispanicum. / Orobanche hederae Vauch. / Hab. in radicibus Hederae Helicis in pomerio / prope Aranjuez / Altitudo: 2500' / ... / Legi die 8 mensi Julii 1844 [m. M. Willkomm]". – 2. "UNIVERSIDAD DE CÓRDOBA / COA / Identificación confirmada / O. hederae Duby / A. Pujadas II 2005" (COI 00056889 - http://coicatalogue.uc.pt/specimen/56889, rev. Ó. Sánchez Pedraja Jul 2020); Hispaniâ ad Aranjuez in praedio regio !" (Reuter 1847: 28, sub O. Hederae Duby); "Cast. nova (Aranjuez in pomerio, Loefl., Reut., Wk.)" (Willkomm 1870: 625, sub O. Hederae Duby; Colmeiro 1888: 279, sub O. Hederae Dub.); "M" (Foley 2001: 61, sub O. hederae Vaucher ex Duby); "Hortus Regius Matritensis (MA) / Orobanchaceae / Orobanche / España, Madrid: Madrid, Predios del Real Jardín / Botánico, setos de Fatsia, flores amarillo pálido, a veces / con tinte vináceo, 06-jun-2012. Creciendo como parásita / de raices superficiales de Fatsia sp. (Araliaceae), J.L. / Fernández-Alonso, 30222 / det. J.L. Fernández (2012) / (5/8)" (NY 02680309; rev. Ó. Sánchez Pedraja Mar 2017); "San Lorenzo de El Escorial (M) 17.6.2015. Inflorescencia. Parásita p. de Hedera sp." (Grijalbo 2016: 296, photo! and Madrid Distribution Map) Málaga (Ma): "cerca de Antequera (Reut., Colm.)" (Colmeiro 1888: 279, sub O. Hederae Dub.); "(Ma)" (Foley 2001: 61, sub O. hederae Vaucher ex Duby). Navarra / Nafarroa / Navarre (Na): BIO 20125 / [...] / Orobanche hederae Duby / Navarra: Villava, confluencia ríos Arga y Ulzama / 30TXN1442, 420 m, 23-jul.-1991 / Aliseda / Leg. I. Biurrum / Det. I. Biurrum // Revisión para «Orobanchaceae - Paleártico» / O. hederae Vaucher ex Duby, Bot. Gall.: 350 (1828) / Det.-rev. Ó. Sánchez Pedraja, 22.09.2006 (Biurrum 1999: 271); "Na" (Foley 2001: 61, sub O. hederae Vaucher ex Duby). Orense / Ourense (Or): "Ourense: Manzaneda, en la orilla izquierda del río Bibei frente a la localidad de As Ermitas (O Bolo), 02-07-1983, F. Gómez Vigide (GV 5928), rev. A.J. Pujadas." (Gómez Vigide & al. 2006: 58, sub O. hederae Vaucher ex Duby). Rioja, La [formerly known as Logroño Province] (Lo): "Lo" (Foley 2001: 61, sub O. hederae Vaucher ex Duby); [en el paquete de Orobanche minor] 1. "Herbario Alejandre / N.º 1240/99 / La Rioja: Viguera, bco. del Arroyo de Madre, ladera / pedregosa, S-SE, bajo cueva Moratel, carrascal abierto, / calizas, 1100 m / 30TWM3880 / 16 de Julio de 1999 / Leg. J.A. Alejandre / M.J. Escalante". – 2. "Revisión para «Orobanchaceae - Paleártico» / O. hederae Vaucher ex Duby / Det.-rev. Ó. Sánchez Pedraja 4-IX-2002". Salamanca (Sa): Monteamayor del Río,15-X-1983, leg. et det. A. Guillén & E. Rico, rev. M.J.Y. Foley et A. Pujadas (herb. SALA 37360); "Sa" (Foley 2001: 61, sub O. hederae Vaucher ex Duby). Segovia (Sg): Sepúlveda, 26-VI-1983, leg. et det. T. Romero, rev. M.J.Y. Foley et A. Pujadas (SALA 40134); "Sg" (Foley 2001: 61, sub O. hederae Vaucher ex Duby). Sevilla / Seville (Se): Sierra Morena, San Nicolás del Puerto, arroyo Húmedo, 30STH60, 22-VI-1976, E.F. Galiano & al. (MA 465689) (Pujadas & al. 2007). Tarragona (T): Montsià, Amposta, Hort del Bou, 31TBF9510 (Balada & al. 2007); "Tarragona" (O. hederae Duby in Herbari Virtual del Mediterrani Occidental, accessed Jan 2012). Teruel (Te): "Aragon (pr. Castellote [Maestrazgo, Aragon], Losc. Pard.)" (Willkomm 1870: 625, sub O. Hederae Duby; Colmeiro 1888: 279, sub O. Hederae Dub.); "(Te)" (Foley 2001: 61, sub O. hederae Vaucher ex Duby). Valencia / València (V): "Játiva [Xàtiva] (Rouy)" (Colmeiro 1888: 279, sub O. Hederae Dub.); BDBC (accessed Jul 2020); "Valencia" (O. hederae Duby in Herbari Virtual del Mediterrani Occidental, accessed Jan 2012). Valladolid (Va): Aguasal, Fuente del Avellano (Gutiérrez Martín 1908). Vizcaya / Bizkaia / Biscay (Bi): "Cantabr. (...Bilbao, Lge. ...)" (Willkomm 1870: 625, sub O. Hederae Duby; Colmeiro 1888: 279, sub O. Hederae Dub.); "Bilbao, jardines de la Casa de Misericordia, 30TWN0490, 25 m, creciendo junto a Hedera helix y Evonymus japonicus, 04.05.1994, M.A. Domingo" (VIT 18192, sub O. hederae [rev. para «Contribuciones VI» / O. hederae Vaucher ex Duby / rev. Ó. Sánchez Pedraja 4-IX-2002]); BIO 0493 / [...] Dupla 1016-82 / Orobanche loricata Rcbh. / 4. Trucios (Bi) Godron VN7992 / Eneka ondoko laso nahasia? / 250 m / 27.05.1982 / C. Aseginolaza // Revisión para «Orobanchaceae - Paleártico» / O. hederae Vaucher ex Duby, Bot. Gall.: 350 (1828) / Det.-rev. Ó. Sánchez Pedraja 22.09.2006 [obs. inflorescencias laxas y largas, más de la mitad del total; corolas contreñidas hacia la garganta]. Zamora (Za): Galende (photo! by Patricio Bariego). Zaragoza (Z): "Z" (Foley 2001: 61, sub O. hederae Vaucher ex Duby). Sweden / Suecia / Suède / Schweden [naturalized] (Pusch 2009: 64 Abb. 80, "synanthrop", sub O. hederae Duby) Scania Province: "Landskrona, gamla kyrkogården, Landskrona s:n, Sk / Nära församlingshemmet, snyltar på planterad murgröna" (https://www.artportalen.se/Image/1848549 - https://www.gbif.org/es/occurrence/1669985078 by Lars Erik Norbäck in Artportalen (Swedish Species Observation System); accessed Aug 2023). Switzerland / Schweiz / Suisse / Svizzera / Suiza (Reichenbach 1829: 42, sub O. medicaginis sensu Rchb.; Reuter 1847: 28, sub O. Hederae Duby; Reichenbach fil. 1862: 103, sub O. Hederae Duby; Beck 1890: 261, sub O. Hederae Duby; Beck 1930: 220, sub O. Hederae Duby; Pusch 2009: 64, sub O. hederae Duby) "Habitat in muris arcis Genevensis [Genève / Genève] ... Vaucher; in muro vetusto ad viam regiam prope Viviscum [Vevey, Vaud / Waadt] Gaudin" (Reichenbach 1829: 42, loc. class. of O. medicaginis sensu Rchb.); "Helvetiâ prope Neuveville (Gibollet !). Neocomum (Godet ! ) et Vevay, (Burnat !)" (Reuter 1847: 28, sub O. Hederae Duby); "Helvetia: prope Basel, Vevey, Lausanne, Genf" (Beck 1890: 261, sub O. Hederae Duby). Turkey / Türkiye / Turquía / Турция [see Asiatic Turkey] Ukraine / Ucrania / Україна / Украина [alien in Ukrainian mainland] Odesa Oblast / Одеська область: "Одеса / Odessa / Odesa [46°26'26.0"N 30°46'04.2"E 46.440547, 30.767842, Odesa Raion / Одеський район], 25 Jun 2023" (https://www.inaturalist.org/observations/169350346 by Anna Petrosyan in iNaturalist; accessed Jul 2020 [yellow form, alien]). Krim / Crimea / Крымский полуостров / Кримський півострів / Tauria / Tauris / Ταυρική [Autonomous Republic of Crimea / Автономная Республика Крым - Республика Крым / Republic of Crimea] (formerly Crimean ASSR / Крымская АССР) [see Ukraine / Russia] (Beck 1890: 261, sub O. Hederae Duby; Beck 1930: 221, sub O. Hederae Duby; Novopokrovskij & Tzvelev 1958: 98[104], sub O. hederae Duby; Seregin 2008: 221, "1169. !! O. hederae Duby*"; Pusch 2009: 64 Abb. 80, sub O. hederae Duby; Tsymbalyuk & Mosyakin 2013: 724, sub O. hederae Duby) "Tauria: prope Nikita [Никита / Нікіта, Yalta Municipality / Ялтинский городской совет] (leg. Steven)" (Beck 1890: 261, sub O. Hederae Duby); 1. "Herbarium Universitatis Mosquensis / 1889 Крым [leg.] Голенкин, М.И. [Golenkin, M.I.]". – 2. "Orobanche Hederae Duby / det. Novopokrovskij VII 1949" (https://plant.depo.msu.ru/open/public/item/MW0626332 - MW 0626332.jpg; rev. Ó. Sánchez Pedraja Jul 2020); 1. "... Orobanche / Оріанgа [?] ...". – 2. "Herbarium Universitatis Mosquensis / 1907 Крым. / Аноним [Anonymous]". – 3. "Orobanche Hederae Duby / det. Novopokrovskij VII 1949" (https://plant.depo.msu.ru/open/public/item/MW0626331 - MW 0626331.jpg; rev. Ó. Sánchez Pedraja Jul 2020); "HERBARIUM UNIVERSITATIS AMICITAE POPULORUM / nomine PATRISII LUMUMBII / Orobanche hederae Duby / Крым / Никитский Ботанический Сад [Nikitsky Botanical Garden (YALT)], Никита / Нікіта, Yalta Municipality / Ялтинский городской совет] / 13.VI.1962. [leg.] Павлова М. [Pavlova M.]" (https://plant.depo.msu.ru/open/public/item/MW0626330 - MW 0626330.jpg; rev. Ó. Sánchez Pedraja Jul 2020); "Herbarium Universitatis Mosquensis (MW) / Orobanche hederae Duby / 44º30'00''N 33º46'50''E 250 м н. у. м. / Крым [Crimea], Балаклавский район [Balaklava District], / 3,5 км к ЗЮЗ от с. Передовое, / днище Чернореченского каньона при выходе из / Байдарской долины, / левый берег р. Черная, / грабинниковый лес с плющом в нижней части / склона каньона, под замшелой известняковой / глыбой. / Собр. Серегин А., Привалова И. [Leg. Seregin A., Privalova I.], 13.09.2003. № T-550 / Atlas florae europaeae: 36-T-WQ4-62" (https://plant.depo.msu.ru/open/public/item/MW0626329 - MW 0626329.jpg; rev. Ó. Sánchez Pedraja Jul 2020); "Herbarium / Moscow State University / Orobanche hederae Duby / Крым, окрестности с. Орлиное [Crimea, the vicinity of village Orlyne (), вдоль дороги, на [along the road, on] / 44°24'11.1"N 33°47'55.3"E / 1.06.2015 / Собр. Фомичев К.И. [Fomichev K.I.]" (https://plant.depo.msu.ru/open/public/item/MW0631561 - MW 0631561.jpg; rev. Ó. Sánchez Pedraja Jul 2020); "1. Крым, Алупка [Alupka], парк. 9 VI 1913. М. Котов [M. Kotov] (KW). 2. Крым, Никитский бот. сад [Nikitsky Botanical Garden (YALT)]. Паразитирует на Hedera taurica. 28 VII 1955. М. Котов та ін. [M. Kotov et al.] (KW)" (Tsymbalyuk & Mosyakin 2013: 724, sub O. hederae Duby); "Балаклавский район, [Balaklava, Sebastopol, 44°35'43.2"N 33°36'38.6"E], 21 Jun 2020" (https://www.inaturalist.org/observations/50441649 - https://www.gbif.org/es/occurrence/2802733246 by Ekaterina Kashirina in iNaturalist; accessed Jul 2020). United Kingdom / Reino Unido / Royaume-Uni / Vereinigtes Königreich (Des Moulins 1835: 80, sub O. hederae; Beck 1890: 261, sub O. Hederae Duby; Beck 1930: 220, sub O. Hederae Duby; Rumsey & Jury 1991: 272-273, sub O. hederae Duby; map of Orobanche hederae [NBNSYS0000004180] in NBN, accessed Jul 2020; map by BSBI, accessed Jul 2020; O. hederae in ukwildflowers.com, accessed Jul 2020; http://herbariaunited.org/specimen/273998 in Herbaria@home - Herbaria United (UK and Irish Herbaria On-line), accessed Jun 2022) Bailiwick of Guernsey: "M. Gay en a encore trouvé quelques échantillons frais, le 10 août 1832, à l'ombre, dans l'île de Guernesey" (Des Moulins 1835: 80, sub O. hederae); "irst discovered in the British Isles on |
