Accepted Name - Synonyms - Ind. loc. [Indicatio locotypica (type locality)] - Lectotypus [Lectotype] - Designated by - Host - Comments - Chromosomatic number - Ic. [Images] - Distribution - Specimens Examined - References - Authors - Cite us ISSN: 2386-9666 - Last Updated: 13 September 2021 |
Accepted Name: Phelypaea tournefortii Desf. in Ann. Mus. Par. 10: 298[-300], t. 21 (1807) (Desfontaines 1807: 298-300, t. 21; 1808, 16, t. 10; Sprengel 1825: 818; Don 1838: 633; Piwowarczyk & al. 2019: 18) Synonyms: Homotypic synonyms: ≡ Phelypaea Orientalis, flore coccineo Tournef., Corollarium: 47 t. 479 (1703) [pre-Linnaean name, not validly published] (Desfontaines 1807: 298, "P. Orientalis flore coccineo. Tournef., Cor. Inst. 47.— Vélins du Muéum"; Desfontaines 1808: 16, "P. Orientalis flore coccineo. Tournef., Cor. Inst. 47.— Vélins du Muéum"; Don 1838: 633; Stapf 1915: 285; Cullen 2010: 380) ≡ Lathraea phelypaea var. β [unnamed] L., Sp. Pl. 2: 606 (1753 [1 May 1753]) ["β Orobanche orientalis, flore coccineo. Tournef. cor. 47."], nom. illeg. (Stapf 1915: 286) ≡ Anoplon tournefortii (Desf.) G. Don, Gen. Hist. 4: 633 (1837-1838) (Piwowarczyk & al. 2019: 18) ≡ Anoplanthus tournefortii (Desf.) Reut. in DC., Prodr. 11: 42 (1847) ["A. Tournefortii Walp. rep. bot. 3 p. 48, ex parte"]; [Walp., Repert. Bot. Syst. 3: 481 (1844 [12-14 Sep 1844]) (Beck 1930: 44; Piwowarczyk & al. 2019: 18)]. Obs. "basi vaginis elongatis inaequalibus involuti. Calyx ... villosus ... (v. s. in h. Bentham.)" (Reuter 1847: 43) ≡ Aeginetia tournefortii (Desf.) [Juss. sec. Steudel] Beck in Engl., Pflanzenr. 96: 44 (1930), pro syn. ≡ Diphelypaea tournefortii (Desf.) Nicolson in Taxon, 24(5-6): 654 (1975 [Nov 1975]), nom. illeg. (Piwowarczyk & al. 2019: 18) Heterotypic synonyms: ? = Anblatum orientale, flore purpurascente Tournef., Corollarium: 48 (1703) [pre-Linnaean name, not validly published]. Syn.: ≡ Lathraea anblatum L., Sp. Pl. 2: 606 (1753 [1 May 1753]) (Linnaeus 1753: 606; Don 1838: 635); = Phelypaea foliata Lamb. in in Trans. Linn. Soc. London 10: 260, t. 7 (1811) (Endlicher 1839: 727) ? = Lathraea anblatum L., Sp. Pl. 2: 606 (1753 [1 May 1753]). Syn.: ≡ Anblatum orientale, flore purpurascente Tournef., Corollarium: 48 (1703) [pre-Linnaean name] (Linnaeus 1753: 606; Don 1838: 635); ≡ Anblatum tournefortii G. Don, Gen. Hist. 4: 635 (1837-1838); = Phelypaea foliata Lamb. in in Trans. Linn. Soc. London 10: 260, t. 7 (1811). Ind. loc.: "Habitat in Oriente" (Linnaeus 1753: 606); "Asia minor" (Sprengel 1825: 818); "H. Native of the Levant" (Don 1838: 635; Lathraea anblatum in the Linnaean Typification Project, accessed Mar 2012). Obs. According to Endlicher (1839: 727), Anblatum orientale, flore purpurascente Tournef., Corollarium: 48 (1703) is conspecific with Phelypaea foliata Lamb. in in Trans. Linn. Soc. London 10: 260, t. 7 (1811). According to Reuter (1847: 43), Anblatum tournefortii G. Don is conspecific with Phelypaea tournefortii Desf. in Ann. Mus. Par. 10: 298. t. 21 (1807). According to Index Synonymique de la Flore de France, Flora iberica and The Plant List, Anblatum tournefortii G. Don is conspecific with Lathraea squamaria L. (1753) [this synonym seems correct on the base of Don's description] ? = Anblatum tournefortii G. Don, Gen. Hist. 4: 635 (1837-1838) (Reuter 1847: 43). Obs. According to Index Synonymique de la Flore de France, Flora iberica and The Plant List, Anblatum tournefortii G. Don is conspecific with Lathraea squamaria L. (1753) ? = Aeginetia coccinea Juss. ex Reut. in DC., Prodr. 11: 44 (1847), pro syn. Syn.: = Anoplanthus tournefortii (Desf.) Reut. in DC., Prodr. 11: 44 (1847) (Reuter 1847: 44) = Anoplanthus coccineus var. peduncularis Boiss., Fl. Orient. 4(2): 494 (1879 [Apr-May 1879]) ["A. coccineus β peduncularis"] (Piwowarczyk & al. 2019: 18). Syn.: = Phelypaea orientalis, flore coccinea Tournef., Corollarium: 47 t. 479 (1703); = Phelypaea tournefortii Desf. in Ann. Mus. Par. 10: 298, t. 21 (1807); = Anoplanthus tournefortii (Desf.) Reut. in DC., Prodr. 11: 42 (1847) (Boissier 1879: 494; Beck 1930: 44; Novopokrovskij & Tzvelev 1958[2000]: 30[28]). Ind. loc.: "in Armeniâ Turcicâ circa Erzerum [Erzurum / Karin, Anatolia, Turkey] (Tourn!, Huet!, Calv!), in Kurdistaniâ (Barré de Lancy [vice-consul de France ŕ Mersine et Adana]! [see P. boissieri], Armeniâ Rossicá (Buhse!), in pratis montosis Georgiae Caucasicae (Hoh!)". Ic.: "Phelypaea Tournefortii Desf. Cor. tab. 10" (Boissier 1879: 494). Obs. "Squamae ad basin caulis superne longe nudis magis congestae" (Boissier 1879: 494) |
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Ind. loc.: "Il est originaire d'Arménie [Western Armenia]" (Desfontaines 1807: 300); "ad radices Pyrethri myriophylli C.A. Meyer [Tanacetum polycephalum subsp. duderanum (Boiss.) Podlech] et Achilleae spec. ... Armenien: Provinz Erzerum [Erzurum, Anatolia, Turkey] (Tournefort..." (Beck 1930: 44; Novopokrovskij & Tzvelev 1958[2000]: 30[28]; Nicolson 1975: 652 "variety B [β]"; Cullen 2010: 380; view IPNI entry). Typus / Type: "Type:—Turkey (lectotype designated here):—Phelypaea Orientalis, flore coccineo [Arménie, nowadays Erzurum Province, Turkey], Tournefort (1703: t. 479!). Turkey (epitype designated here):—Armenia, im valle Kapuclu, inter Baiburt [Bayburt] et Erzeroum [Erzurum], Maio 1853, Huet de Pavillon [Plantae Orientales exsiccatae] (P-02970941!)." (Piwowarczyk & al. 2019: 18, sub D. tournefortii). N.B. "that no specimens of the Phelipaea Tournefortii now remain in his Herbarium, or are known to exist in any other collection" (Lambert 1811: 261); "No specimen of this species could be found in the Tournefort herbarium (P)" (Gilli 1982: 2); "There is, apparently, no specimen in the Tournefort herbarium, but Aubriet, the artist who accompanied Tournefort, made an illustration which appears as t. 479 of the Corollarium, and can be regarded as the type material; it is clearly the plant recognised here as Phelypaea tournefortii." (Cullen 2010: 380; Piwowarczyk & al. 2019: 18, sub D. tournefortii). Designated by: Piwowarczyk & al. in Phytotaxa 386(1): 18 (2019). Host: Parasitic exclusively on Tanacetum species (Asteraceae / Compositae) (Annotated Checklist of Host Plants of Orobanchaceae, accessed Aug 2009); "ad radices Pyrethri myriophylli C.A. Meyer [Tanacetum polycephalum subsp. duderanum (Boiss.) Podlech [non Tanacetum myriophyllum Willd., nom. illeg.]] et Achilleae spec. [Tanacetum sp. ?]" (Reuter 1847: 43; Fischer 1852: 107, [Armenié, Bushe], sub Anoplanthus Tournefortii; Beck 1930: 44; Novopokrovskij & Tzvelev 1958[2000]: 30[28]); "On Achillea and Tanacetum" (Gilli 1982: 2); "Tanacetum sp. (Asteraceae), mainly T. polycephalum subsp. argyrophyllum (K. Koch) Podlech [syn. T. argyrophyllum (C. Koch) Tzvel.], T. chiliophyllum (Fisch. & C. A. Mey.) Sch. Bip., T. canescens DC. [Pyrethrum canescens (DC.) Boiss.]. Need confirmation: Anthemis L., "Pyrethrum myriophyllum (Willd.) C.A.M." [Tanacetum polycephalum subsp. duderanum (Boiss.) Podlech] (Novopokrovskij & Tzvelev 1958)." (Piwowarczyk & al. 2019: 19, sub D. tournefortii). Tanacetum aureum (Lam.) Greuter, M. V. Agab. & Wagenitz [Tanacetum chiliophyllum (Fisch. & C.A. Mey.) Sch. Bip.; Achillea aurea Lam.; Pyrethrum chiliophyllum DC.] (Piwowarczyk & al. 2019: 19, Armenia, sub D. tournefortii). Tanacetum canescens DC. [Pyrethrum canescens (DC.) Boiss.] (Piwowarczyk & al. 2019: 19, Armenia, sub D. tournefortii). Tanacetum polycephalum subsp. argyrophyllum (K. Koch) Podlech [T. argyrophyllum (C. Koch) Tzvel.; Tanacetum myriophyllum Willd., nom. illeg.; Achillea bipinnata L.; Pyrethrum myriophyllum auct. plur.] (Piwowarczyk & al. 2019: 19, Armenia, sub D. tournefortii). Tanacetum polycephalum subsp. duderanum (Boiss.) Podlech [Pyrethrum duderanum Boiss.; Pyrethrum myriophyllum sensu C.A. Mey. [non Tanacetum myriophyllum Willd., nom. illeg.]] (Fischer 1852: 107, [Nakhchivan (Azerbaijan), Bushe], sub Anoplanthus Tournefortii; Stapf 1915: 293, [Nakhchivan (Azerbaijan), Bushe], sub P. tournefortii; Beck 1930: 44, sub P. tournefortii; Novopokrovskij & Tzvelev 1958[2000]: 30[28]). Comments: Morphological observations: "P. Scapo basi involuto, villoso ... Calice ... velues ... Le Phelipaea de Tournefort a les tiges nues et les divisiones de la corolle sont circulaires et non ovales" (Desfontaines 1807: 298-300). Key to these species (Stapf 1915: 290; Novopokrovskii & Tzvelev 1958[2000]: 28[26]; Cullen 2010: 383-384); pollen morphology (Zare & al. 2014); key (Piwowarczyk & al. 2019: 19). Phylogenetic observations: Carlón & al. in Documentos Jard. Bot. Atlántico (Gijón) 3: 57 [x = 19, sub Phelypaea (syn. Diphelypaea)], 61 f. 4 ["Phylogenetic relationships", sub Phelypaea], 63 (2005); "Recent molecular phylogenetic studies suggest two major lineages, informally called the Orobanche-group and the Phelipanche-group (Schneeweiss et al., 2004a). The first includes Orobanche sect. Orobanche and the small SW Asian genus Phelypaea (listed as Diphelypaea in Schneeweiss et al., 2004a), while the second includes the remaining sections of Orobanche (sects. Gymnocaulis, Myzorrhiza, and Trionychon). The circumscription of these groups is well supported by the distribution of chromosome base numbers, these being x = 19 in the Orobanche-group and x = 12 in the Phelipanche-group (Schneeweiss et al. 2004b)" Weiss-Schneeweiss & al., 2006; Schneider, 2016: f. 1. Phelypaea L., Op. Var.: 237 (1758) is the correct name for the genus, because the previous names were not validly published according to ICN (Turland & al. 2018). Phelypaea Hill in Scott, Suppl. Chambers Cyclop. (1753) [not validly published] (Lonchamp in INRA-MNHN 1999; HUH id=16687, accessed Jan 2020; Turland & al. 2018: Art. 34, "suppressed work" (see App. I)). Type Name: Lathraea phelypaea L., Sp. Pl.: 606 (1753) Phelypaea Hill in Owen, Dict. Arts Sci. 3: 2416 (1755 [1 Mar 1755]) [not validly published] ["Phelypaea, in botany, a genus of the didynamia-angiospermia..."] (Reveal 1991: 228, "The Hill genus name is now the oldest one available"; Turland 2018: Art. 34, "suppressed work" (see App. I)). NT [Neotype]: Lathraea phelypaea L., Sp. Pl.: 606 (1753) (Reveal 1991: 228); ≡ Cistanche phelypaea (L.) Cout. (1913) Phelypaea P. Browne, Civ. Hist. Jamaica: 269 (1756 [Mar 1756]) [not validly published] (Shenzhen Code - Art. 38.6 Ex. 13, "In publishing "Phaelypea" without a generic description or diagnosis, Browne (Civ. Nat. Hist. Jamaica: 269. 1756) included and described a single species [Phelypaea erecta P. Browne], but he gave the species a phrase name (McNeill & Greuter 2016) not a validly published binomial. Art. 38.5 does not therefore apply and "Phaelypea" is not a validly published name.") (Turland & al. 2018: Art. 38.6 Ex. 13). Phelypaea L., Op. Var.: 237 (1758), non Phelypaea Hill (1755) nec Phelypaea P. Browne (1756) ["Phaelypae"] (Reveal 1991: 228; Nicolson 1975: 653). Type Name: Phelypaea tournefortii Desf. (1807); ≡ Diphelypaea tournefortii (Desf.) Nicolson (1975). Obs. Carlón & al. in Documentos Jard. Bot. Atlántico (Gijón) 3: 57, 58, 61 f. 4, 63 (2005); "vide Dandy, Ind. Gen. Vasc. Pl. 1753-74 (Regn. Veg. li.) 7 (1967)". Replaced synonym of: Diphelypaea Nicolson Taxon 24(5-6): 654. 1975 (view IPNI entry, accessed Feb 2008). "Dr R.K. Brummitt of the Royal Botanic Gardens, Kew tells me that he regards the name Phelypaea as the correct one for the genus." (Cullen 2010: 380). Diphelypaea Nicolson in Taxon, 24(5-6): 654 (1975), nom. nov. [nom. illeg.]. Syn.: ≡ Phelypaea L., Op. Var.: 237 (1758). Type Name: Diphelypaea tournefortii (Desf.) Nicolson (1975) [Phelypaea tournefortii Desf. (1807)]. IPNI: "without full basionym ref.". The reference to the place of publication of the basionym in "References"! (Nicolson 1975). Chromosomatic number: n = 19, 2n = 38 (Schneeweiss & al. 2004, sub Diphelypaea; Carlón & al. 2005: 57, sub Phelypaea [between Bayburt and Erzurum, NE Anatolia, Turkey]). Ic.: a, Tournefort, Corollarium: 47 t. 479 (1703), sub P. orientalis, flore coccinea; b, Desfontaines 1807: t. 21 et 1808: 16, t. 10, sub Phelypaea tournefortii; c, Beck in Engl., Pflanzenr. 96: 43 f. 9 (1930, sub P. Tournefortii); d, habit (Muradiye, Van, Turkey [May 2009, photo by P. Coulot]); e, flower (ibidem); f, calyx (ib. ["Calyx ... villosus" (Reuter 1847: 43)]); g, scale-leaves (ib. ["Squamae ad basin caulis superne longe nudis magis congestae" (Boissier 1879: 494)]); h, habit (P. tournefortii eastern Anatolia, Jul 2002 by G. Schneeweiss); i, flower (P. tournefortii eastern Turkey, by L.J. Musselman); j, sheet (Turkey in Demirkuş 2007); k, Novopokrovskij & Tzvelev (1958[2000]: 30[37] t, 3 f. 5); l, D. tournefortii vs. D. coccinea vs. D. boissieri (P, habits and calyces); n, Piwowarczyk & al. 2019: 18 f. 9 A-F, sub D. tournefortii. Distribution: Eurasian distribution (Armenia, Azerbaijan, Georgia [needs be confirmed], Iran [needs be confirmed], Turkey). Eurasian distribution (Domina & Raab-Straube 2010, sub Phelypaea tournefortii, accessed Jul 2010). |
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Specimens examined and literature records.
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