Accepted Name - Synonyms - Ind. loc. [Indicatio locotypica (type locality)] - Lectotypus [Lectotype] - Designated by - Host - Comments - Chromosomatic number - Ic. [Images] - Distribution - Specimens Examined - References - Authors - Cite us

ISSN: 2386-9666 - Last Updated: 13 September 2021

Accepted Name:

Phelypaea tournefortii Desf. in Ann. Mus. Par. 10: 298[-300], t. 21 (1807) (Desfontaines 1807: 298-300, t. 21; 1808, 16, t. 10; Sprengel 1825: 818; Don 1838: 633; Piwowarczyk & al. 2019: 18)

Synonyms:

Homotypic synonyms:

≡ Phelypaea Orientalis, flore coccineo Tournef., Corollarium: 47 t. 479 (1703) [pre-Linnaean name, not validly published] (Desfontaines 1807: 298, "P. Orientalis flore coccineo. Tournef., Cor. Inst. 47.— Vélins du Muéum"; Desfontaines 1808: 16, "P. Orientalis flore coccineo. Tournef., Cor. Inst. 47.— Vélins du Muéum"; Don 1838: 633; Stapf 1915: 285; Cullen 2010: 380)

≡ Lathraea phelypaea var. β [unnamed] L., Sp. Pl. 2: 606 (1753 [1 May 1753]) ["β Orobanche orientalis, flore coccineo. Tournef. cor. 47."], nom. illeg. (Stapf 1915: 286)

≡ Anoplon tournefortii (Desf.) G. Don, Gen. Hist. 4: 633 (1837-1838) (Piwowarczyk & al. 2019: 18)

≡ Anoplanthus tournefortii (Desf.) Reut. in DC., Prodr. 11: 42 (1847) ["A. Tournefortii Walp. rep. bot. 3 p. 48, ex parte"]; [Walp., Repert. Bot. Syst. 3: 481 (1844 [12-14 Sep 1844]) (Beck 1930: 44; Piwowarczyk & al. 2019: 18)]. Obs. "basi vaginis elongatis inaequalibus involuti. Calyx ... villosus ... (v. s. in h. Bentham.)" (Reuter 1847: 43)

≡ Aeginetia tournefortii (Desf.) [Juss. sec. Steudel] Beck in Engl., Pflanzenr. 96: 44 (1930), pro syn.

≡ Diphelypaea tournefortii (Desf.) Nicolson in Taxon, 24(5-6): 654 (1975 [Nov 1975]), nom. illeg. (Piwowarczyk & al. 2019: 18)

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Heterotypic synonyms:

? = Anblatum orientale, flore purpurascente Tournef., Corollarium: 48 (1703) [pre-Linnaean name, not validly published]. Syn.: ≡ Lathraea anblatum L., Sp. Pl. 2: 606 (1753 [1 May 1753]) (Linnaeus 1753: 606; Don 1838: 635); = Phelypaea foliata Lamb. in in Trans. Linn. Soc. London 10: 260, t. 7 (1811) (Endlicher 1839: 727)

? = Lathraea anblatum L., Sp. Pl. 2: 606 (1753 [1 May 1753]). Syn.: ≡ Anblatum orientale, flore purpurascente Tournef., Corollarium: 48 (1703) [pre-Linnaean name] (Linnaeus 1753: 606; Don 1838: 635); ≡ Anblatum tournefortii G. Don, Gen. Hist. 4: 635 (1837-1838); = Phelypaea foliata Lamb. in in Trans. Linn. Soc. London 10: 260, t. 7 (1811). Ind. loc.: "Habitat in Oriente" (Linnaeus 1753: 606); "Asia minor" (Sprengel 1825: 818); "H. Native of the Levant" (Don 1838: 635; Lathraea anblatum in the Linnaean Typification Project, accessed Mar 2012). Obs. According to Endlicher (1839: 727), Anblatum orientale, flore purpurascente Tournef., Corollarium: 48 (1703) is conspecific with Phelypaea foliata Lamb. in in Trans. Linn. Soc. London 10: 260, t. 7 (1811). According to Reuter (1847: 43), Anblatum tournefortii G. Don is conspecific with Phelypaea tournefortii Desf. in Ann. Mus. Par. 10: 298. t. 21 (1807). According to Index Synonymique de la Flore de France, Flora iberica and The Plant List, Anblatum tournefortii G. Don is conspecific with Lathraea squamaria L. (1753) [this synonym seems correct on the base of Don's description]

? = Anblatum tournefortii G. Don, Gen. Hist. 4: 635 (1837-1838) (Reuter 1847: 43). Obs. According to Index Synonymique de la Flore de France, Flora iberica and The Plant List, Anblatum tournefortii G. Don is conspecific with Lathraea squamaria L. (1753)

? = Aeginetia coccinea Juss. ex Reut. in DC., Prodr. 11: 44 (1847), pro syn. Syn.: = Anoplanthus tournefortii (Desf.) Reut. in DC., Prodr. 11: 44 (1847) (Reuter 1847: 44)

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= Anoplanthus coccineus var. peduncularis Boiss., Fl. Orient. 4(2): 494 (1879 [Apr-May 1879]) ["A. coccineus β peduncularis"] (Piwowarczyk & al. 2019: 18). Syn.: = Phelypaea orientalis, flore coccinea Tournef., Corollarium: 47 t. 479 (1703); = Phelypaea tournefortii Desf. in Ann. Mus. Par. 10: 298, t. 21 (1807); = Anoplanthus tournefortii (Desf.) Reut. in DC., Prodr. 11: 42 (1847) (Boissier 1879: 494; Beck 1930: 44; Novopokrovskij & Tzvelev 1958[2000]: 30[28]). Ind. loc.: "in Armeniâ Turcicâ circa Erzerum [Erzurum / Karin, Anatolia, Turkey] (Tourn!, Huet!, Calv!), in Kurdistaniâ (Barré de Lancy [vice-consul de France ŕ Mersine et Adana]! [see P. boissieri], Armeniâ Rossicá (Buhse!), in pratis montosis Georgiae Caucasicae (Hoh!)". Ic.: "Phelypaea Tournefortii Desf. Cor. tab. 10" (Boissier 1879: 494). Obs. "Squamae ad basin caulis superne longe nudis magis congestae" (Boissier 1879: 494)

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Ind. loc.: "Il est originaire d'Arménie [Western Armenia]" (Desfontaines 1807: 300); "ad radices Pyrethri myriophylli C.A. Meyer [Tanacetum polycephalum subsp. duderanum (Boiss.) Podlech] et Achilleae spec. ... Armenien: Provinz Erzerum [Erzurum, Anatolia, Turkey] (Tournefort..." (Beck 1930: 44; Novopokrovskij & Tzvelev 1958[2000]: 30[28]; Nicolson 1975: 652 "variety B [β]"; Cullen 2010: 380; view IPNI entry).

Typus / Type: "Type:—Turkey (lectotype designated here):—Phelypaea Orientalis, flore coccineo [Arménie, nowadays Erzurum Province, Turkey], Tournefort (1703: t. 479!). Turkey (epitype designated here):—Armenia, im valle Kapuclu, inter Baiburt [Bayburt] et Erzeroum [Erzurum], Maio 1853, Huet de Pavillon [Plantae Orientales exsiccatae] (P-02970941!)." (Piwowarczyk & al. 2019: 18, sub D. tournefortii).

N.B. "that no specimens of the Phelipaea Tournefortii now remain in his Herbarium, or are known to exist in any other collection" (Lambert 1811: 261); "No specimen of this species could be found in the Tournefort herbarium (P)" (Gilli 1982: 2); "There is, apparently, no specimen in the Tournefort herbarium, but Aubriet, the artist who accompanied Tournefort, made an illustration which appears as t. 479 of the Corollarium, and can be regarded as the type material; it is clearly the plant recognised here as Phelypaea tournefortii." (Cullen 2010: 380; Piwowarczyk & al. 2019: 18, sub D. tournefortii).

Designated by: Piwowarczyk & al. in Phytotaxa 386(1): 18 (2019).

Host: Parasitic exclusively on Tanacetum species (Asteraceae / Compositae) (Annotated Checklist of Host Plants of Orobanchaceae, accessed Aug 2009); "ad radices Pyrethri myriophylli C.A. Meyer [Tanacetum polycephalum subsp. duderanum (Boiss.) Podlech [non Tanacetum myriophyllum Willd., nom. illeg.]] et Achilleae spec. [Tanacetum sp. ?]" (Reuter 1847: 43; Fischer 1852: 107, [Armenié, Bushe], sub Anoplanthus Tournefortii; Beck 1930: 44; Novopokrovskij & Tzvelev 1958[2000]: 30[28]); "On Achillea and Tanacetum" (Gilli 1982: 2); "Tanacetum sp. (Asteraceae), mainly T. polycephalum subsp. argyrophyllum (K. Koch) Podlech [syn. T. argyrophyllum (C. Koch) Tzvel.], T. chiliophyllum (Fisch. & C. A. Mey.) Sch. Bip., T. canescens DC. [Pyrethrum canescens (DC.) Boiss.]. Need confirmation: Anthemis L., "Pyrethrum myriophyllum (Willd.) C.A.M." [Tanacetum polycephalum subsp. duderanum (Boiss.) Podlech] (Novopokrovskij & Tzvelev 1958)." (Piwowarczyk & al. 2019: 19, sub D. tournefortii).

Tanacetum aureum (Lam.) Greuter, M. V. Agab. & Wagenitz [Tanacetum chiliophyllum (Fisch. & C.A. Mey.) Sch. Bip.; Achillea aurea Lam.; Pyrethrum chiliophyllum DC.] (Piwowarczyk & al. 2019: 19, Armenia, sub D. tournefortii).

Tanacetum canescens DC. [Pyrethrum canescens (DC.) Boiss.] (Piwowarczyk & al. 2019: 19, Armenia, sub D. tournefortii).

Tanacetum polycephalum subsp. argyrophyllum (K. Koch) Podlech [T. argyrophyllum (C. Koch) Tzvel.; Tanacetum myriophyllum Willd., nom. illeg.; Achillea bipinnata L.; Pyrethrum myriophyllum auct. plur.] (Piwowarczyk & al. 2019: 19, Armenia, sub D. tournefortii).

Tanacetum polycephalum subsp. duderanum (Boiss.) Podlech [Pyrethrum duderanum Boiss.; Pyrethrum myriophyllum sensu C.A. Mey. [non Tanacetum myriophyllum Willd., nom. illeg.]] (Fischer 1852: 107, [Nakhchivan (Azerbaijan), Bushe], sub Anoplanthus Tournefortii; Stapf 1915: 293, [Nakhchivan (Azerbaijan), Bushe], sub P. tournefortii; Beck 1930: 44, sub P. tournefortii; Novopokrovskij & Tzvelev 1958[2000]: 30[28]).

Comments:

Morphological observations: "P. Scapo basi involuto, villoso ... Calice ... velues ... Le Phelipaea de Tournefort a les tiges nues et les divisiones de la corolle sont circulaires et non ovales" (Desfontaines 1807: 298-300). Key to these species (Stapf 1915: 290; Novopokrovskii & Tzvelev 1958[2000]: 28[26]; Cullen 2010: 383-384); pollen morphology (Zare & al. 2014); key (Piwowarczyk & al. 2019: 19).

Phylogenetic observations: Carlón & al. in Documentos Jard. Bot. Atlántico (Gijón) 3: 57 [x = 19, sub Phelypaea (syn. Diphelypaea)], 61 f. 4 ["Phylogenetic relationships", sub Phelypaea], 63 (2005); "Recent molecular phylogenetic studies suggest two major lineages, informally called the Orobanche-group and the Phelipanche-group (Schneeweiss et al., 2004a). The first includes Orobanche sect. Orobanche and the small SW Asian genus Phelypaea (listed as Diphelypaea in Schneeweiss et al., 2004a), while the second includes the remaining sections of Orobanche (sects. Gymnocaulis, Myzorrhiza, and Trionychon). The circumscription of these groups is well supported by the distribution of chromosome base numbers, these being x = 19 in the Orobanche-group and x = 12 in the Phelipanche-group (Schneeweiss et al. 2004b)" Weiss-Schneeweiss & al., 2006; Schneider, 2016: f. 1.

Taxonomic observations:

Phelypaea L., Op. Var.: 237 (1758) is the correct name for the genus, because the previous names were not validly published according to ICN (Turland & al. 2018).

Phelypaea Hill in Scott, Suppl. Chambers Cyclop. (1753) [not validly published] (Lonchamp in INRA-MNHN 1999; HUH id=16687, accessed Jan 2020; Turland & al. 2018: Art. 34, "suppressed work" (see App. I)). Type Name: Lathraea phelypaea L., Sp. Pl.: 606 (1753)

Phelypaea Hill in Owen, Dict. Arts Sci. 3: 2416 (1755 [1 Mar 1755]) [not validly published] ["Phelypaea, in botany, a genus of the didynamia-angiospermia..."] (Reveal 1991: 228, "The Hill genus name is now the oldest one available"; Turland 2018: Art. 34, "suppressed work" (see App. I)). NT [Neotype]: Lathraea phelypaea L., Sp. Pl.: 606 (1753) (Reveal 1991: 228); ≡ Cistanche phelypaea (L.) Cout. (1913)

Phelypaea P. Browne, Civ. Hist. Jamaica: 269 (1756 [Mar 1756]) [not validly published] (Shenzhen Code - Art. 38.6 Ex. 13, "In publishing "Phaelypea" without a generic description or diagnosis, Browne (Civ. Nat. Hist. Jamaica: 269. 1756) included and described a single species [Phelypaea erecta P. Browne], but he gave the species a phrase name (McNeill & Greuter 2016) not a validly published binomial. Art. 38.5 does not therefore apply and "Phaelypea" is not a validly published name.") (Turland & al. 2018: Art. 38.6 Ex. 13).

Phelypaea L., Op. Var.: 237 (1758), non Phelypaea Hill (1755) nec Phelypaea P. Browne (1756) ["Phaelypae"] (Reveal 1991: 228; Nicolson 1975: 653). Type Name: Phelypaea tournefortii Desf. (1807); ≡ Diphelypaea tournefortii (Desf.) Nicolson (1975). Obs. Carlón & al. in Documentos Jard. Bot. Atlántico (Gijón) 3: 57, 58, 61 f. 4, 63 (2005); "vide Dandy, Ind. Gen. Vasc. Pl. 1753-74 (Regn. Veg. li.) 7 (1967)". Replaced synonym of: Diphelypaea Nicolson Taxon 24(5-6): 654. 1975 (view IPNI entry, accessed Feb 2008). "Dr R.K. Brummitt of the Royal Botanic Gardens, Kew tells me that he regards the name Phelypaea as the correct one for the genus." (Cullen 2010: 380).

Diphelypaea Nicolson in Taxon, 24(5-6): 654 (1975), nom. nov. [nom. illeg.]. Syn.: ≡ Phelypaea L., Op. Var.: 237 (1758). Type Name: Diphelypaea tournefortii (Desf.) Nicolson (1975) [Phelypaea tournefortii Desf. (1807)]. IPNI: "without full basionym ref.". The reference to the place of publication of the basionym in "References"! (Nicolson 1975).

Chromosomatic number: n = 19, 2n = 38 (Schneeweiss & al. 2004, sub Diphelypaea; Carlón & al. 2005: 57, sub Phelypaea [between Bayburt and Erzurum, NE Anatolia, Turkey]).

Ic.: a, Tournefort, Corollarium: 47 t. 479 (1703), sub P. orientalis, flore coccinea; b, Desfontaines 1807: t. 21 et 1808: 16, t. 10, sub Phelypaea tournefortii; c, Beck in Engl., Pflanzenr. 96: 43 f. 9 (1930, sub P. Tournefortii); d, habit (Muradiye, Van, Turkey [May 2009, photo by P. Coulot]); e, flower (ibidem); f, calyx (ib. ["Calyx ... villosus" (Reuter 1847: 43)]); g, scale-leaves (ib. ["Squamae ad basin caulis superne longe nudis magis congestae" (Boissier 1879: 494)]); h, habit (P. tournefortii eastern Anatolia, Jul 2002 by G. Schneeweiss); i, flower (P. tournefortii eastern Turkey, by L.J. Musselman); j, sheet (Turkey in Demirkuş 2007); k, Novopokrovskij & Tzvelev (1958[2000]: 30[37] t, 3 f. 5); l, D. tournefortii vs. D. coccinea vs. D. boissieri (P, habits and calyces); n, Piwowarczyk & al. 2019: 18 f. 9 A-F, sub D. tournefortii.

Distribution: Eurasian distribution (Armenia, Azerbaijan, Georgia [needs be confirmed], Iran [needs be confirmed], Turkey).

Eurasian distribution (Domina & Raab-Straube 2010, sub Phelypaea tournefortii, accessed Jul 2010).

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Specimens examined and literature records.

Armenia / Hayastan / Armenien / Arménie / Армения (Stapf 1915: 293, sub Anoplanthus coccineus var. nigrovittatus; Beck 1930: 44, sub P. tournefortii; Piwowarczyk & al. 2019: 19-21, f. 10, sub D. tournefortii)

Ararat Province / Араратская область: "Herbarium Universitatis Mosquеnsis (MW) / Phelураea tournеfortii Desf. / Армения. / Вединский район [Vedi rayonu / Ararat rayonu] / В 2 км к югу от с. Кярки [2 km to the south of the village of Karki / Kərki / Kyarki / Tigranashen / Тигранашен (Садаракский район / Sadarak District)] можжевеловое редколесье, 1400 м. / 29 мая 1973 г. [29.05.1973] Н. Шведчикова [N. Shvedchikova]" (https://plant.depo.msu.ru/open/public/item/MW0723306 - MW 0723306.jpg - https://www.gbif.org/occurrence/1697840797; rev. Ó. Sánchez Pedraja Apr 2018, on Tanacetum sp.); "Herbarium Universitatis Mosquеnsis (MW) / Phelураea tournеfortii Desf. / Армения. / Вединский район [Vedi rayonu / Ararat rayonu] / Окрестности Советаmена [Surroundings of the Council (Vedi)]. / Н. Шведчикова [N. Shvedchikova] 30 мая 1973 [30.05.1973] г. 7782" (https://plant.depo.msu.ru/open/public/item/MW0723307 - MW 0723307.jpg - https://www.gbif.org/occurrence/1697840788; rev. Ó. Sánchez Pedraja Apr 2018); "Plants of Armenia / ... / Flile NY: 288.0 / Phelipaea coccinea (M. Bieb.) Poir. / det. K. Tamanyan, / Armenia. 3 km S from Lusashoh village [Lusashogh / Garaxach / Лусашох], 39ş49'72N, 44ş59'195E / 1760 m. (... / ...). Mountain steppe. / Kamilla Tamanyan 10-2005  28 May 2005 / George Fayvush / The New York Botanical Garden (NY) / in collaboration with / Institute of Botany, Armenian Academy of Sciences (ERE)" (https://science.mnhn.fr/institution/mnhn/collection/p/item/p03931901 - P 03931901.jpg - https://www.gbif.org/occurrence/607811327; rev. Ó. Sánchez Pedraja Apr 2018); "Цветущее растение. Армения, Гарни, Хосровский заповедник [Garni, Khosrov Reserva] 18.05.2010" (http://www.plantarium.ru/page/image/id/107519.html by Ваэ Мартиросян [Vahe Martirosyan], 2011 in Plantarium [Diphelypaea tournefortii, on Tanacetum sp.] and iNaturalist; accessed Feb 2013); "Khosrov reserve 2295 m a.s.l.", 5 June 2007 (https://www.inaturalist.org/observations/585400 - http://www.inaturalist.org/photos/744571 by Roberto Sindaco, sub Diphelypaea, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Jan 2015 [the photo seems to correspond to P. tournefortii on Tanacetum sp.).

Gegharkunik Province / Гехаркуникская область: "im Distr. Novobajazet [Novo Bajazet / Nor Bayezet / Novyi Bayazet, Ajridzha / Armaghan Lerr] um Solak, Berezovaja butka, Konstantinowska (E. Mylow!), bei Station Eilar [Dzak] (A. Fomin!)" (Beck 1930: 44, sub P. tournefortii); "Diphelypaea tournefortii, filmed in the Tarachichaki [Tsaghkunk / Varazhnunik] (town) ..." (http://ic.pics.livejournal.com/vahemart/27418323/85148/85148_original.jpg by Ваэ Мартиросян [Vahe Martirosyan], 2011 in vahemart.livejournal.com; accessed Jan 2015).

Kotayk / Kotaik Province / Котайкская область: "Herbarium Universitatis Mosquеnsis / Plantae Caucasicae Collaboratoribus Horti / Botanici Universitatis Mosquеnsis Lectaе / Phelураea tournеfortii Desf. / Армения, Разданский р-н [Hrazdan raion], / хр. Цахкунец [Tsaghkunyats Mountains], басс. р. Мармарик [Marmarik], / окр. пос. Цахкадзор [Tsaghkadzor / Tsakhkadzor] / М.Г. Пименов Е.В. Клюйков / Л.П. Томкович Н.А. Кириллина [M.G. Pimenov, E.V. Klyuykov / L.P. Tomkovich, N.A. Cyrillin] / 5 июня 1977 [5 June 1977]" (https://plant.depo.msu.ru/open/public/item/MW0723305 - MW 0723305.jpg - https://www.gbif.org/occurrence/1697840793; rev. Ó. Sánchez Pedraja Apr 2018, on Tanacetum sp.); Armenia: Kotayk province, Nairi distr., c. 3 km N Yeghvard, mountain Arailer [Ara / Араилер], "Tsahhkevank", 2002-06-29, leg. G. Fayvush s.n. OPTIMA Iter XI to Armenia Rec. It 2373 (B 10 0469878); "Diphelypaea tournefortii, Berdatak, 2010.05.18 // Tsaghkadzor [Tsakhkadzor / Darachichag]" (photos 5788747528409755042 and 179759 [https://www.inaturalist.org/observations/80313] by Vahe Martirosyan [Ваэ Мартиросян] in Flora of Armenia's Check List and iNaturalist.org; accessed Feb 2013); "Цветок. Армения, область Котайк, окрестности Цахкадзора [neighborhood Tsakhkadzor] 31.05.2008" (photo 107214 by Ваэ Мартиросян [Vahe Martirosyan], 2011 in Plantarium [Diphelypaea tournefortii]; accessed Feb 2013); "Kotayk' [Yelgovan], Armenia" (http://static.panoramio.com/photos/original/57543824.jpg by Vahe Martirosyan in www.panoramio.com, accessed Mar 2012); "Цветущее растение. Армения, обл. Котайк, окр. села Вохчаберд [env. Voghjaberd village], разнотравная степь [grassland steppe]. 26.05.2012" (http://www.plantarium.ru/page/image/id/166927.html et 166926 by Владислав Куропаткин [Vladislav Kuropatkin] in Plantarium [Diphelypaea tournefortii]; accessed Feb 2013); "Diphelypaea tournefortii, Tsaghkadzor, 2014.06.04" (photo 6024440769631112354 by Vahe Martirosyan [Ваэ Мартиросян] in http://vahemart.livejournal.com; accessed Jan 2015); "Photo taken in Jrvezh Forest Park, H-3, Jrvezh, Armenia 40° 10' 35.44" N  44° 37' 51.03" E" (http://static.panoramio.com/photos/large/3909676.jpg by in www.panoramio.com; accessed Jan 2015); "Armenia, near Tseghkadrov city, dry grassland 40,524303ş N ; 44722640ş E. 1 5.05.2014. M. Peregrym" (Леонідович [Leonidovich] 2016: 106, sub P. tournefortii).

Vayots Dzor Province / Вайоцдзорская область: "Herbarium Universitatis Mosquеnsis (MW) / Phelураea tournеfortii Desf. / Армянская ССР, Даралaгез [Armenian SSR, Daralagez]. Гора Теке-Долдуран, урочище Xазма [T’ek’sari Lerrnashght’a range?, natural boundary Hazma?], / горные луг [mountain meadow], 10 июля 1929 г. [10.07.1929] / Собрали Ант. и И.llyкины [Leg. Ant. and I. llykin]" (https://plant.depo.msu.ru/open/public/item/MW0723304 - MW 0723304.jpg - https://www.gbif.org/occurrence/1697840810; rev. Ó. Sánchez Pedraja Apr 2018); "... / Diphelypaea tournefortii ... / ... Armenia, Vayotsdzor province, Vayk distr., side of volcano Vayots Sar, loc. Topasi / Dalik, 4 km SWW of village Karmrashen, 2400 m.s.m.; 39ş48' / 45ş29'E / Leg. M. Barkworth, F. Smith, E. Gabrielian, A. Nersesyan, / 27.06.2003 / Dt. M. Oganesian 2003" (UTC 00239126 - https://s.idigbio.org/idigbio-images-prod/3a60a4764186884f15b7485bbeabe6d3 - https://www.gbif.org/es/occurrence/1986951694; rev. Ó. Sánchez Pedraja Feb 2020); Proshaberd [39°49'54.1"N 45°22'27.0"E, near Vernashen, Gladzor Municipality], 18 Jun 2012 (https://www.inaturalist.org/observations/23155801 - https://www.gbif.org/occurrence/2823126326 by Christoph Schmidtlein, sub Phelypaea coccinea, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Jun 2021 [the photos correspond to P. tournefortii on Tanacetum sp.); "Цветки. Армения, Селимский перевал [Selim Pass / Orbelian's Caravanserai]. 20.05.2014" (http://www.plantarium.ru/page/image/id/238873.html by Тамара Галстян [Tamara Galstyan] 2014 in Plantarium [Diphelypaea tournefortii]; accessed Jan 2015); "Цветок. Армения, Вайоц Дзор, окр. монастыря Спитакавор [Vayots Dzor, env. Spitakavor monastery]. 02.05.2013" (http://www.plantarium.ru/page/image/id/188448.html et http://www.plantarium.ru/page/image/id/188447.html by Марина Скотникова [Marina Skotnikova] 2013 in Plantarium [Diphelypaea tournefortii]; accessed Jan 2015).

Yerevan / Erywan / Jerewan / Ереван: "Caucasus, Gouv. Eriwan, St. Eyliar, on Pyrethrum myriophyllum, Thomin (in Acta Hort. Tifl. vi. II. (1902) 56)" (Stapf 1915: 293, sub Anoplanthus coccineus var. nigrovittatus) [Pyrethrum myriophyllum is the typical host of D. tournefortii]; "Distr. Eriwan [Erywan / Yerevan / Jerewan / Ереван]: bei Hilar, zwischen Garyna und Arabat (Bayern)" (Beck 1930: 44, sub P. tournefortii).

Unknown location: "Herbarium Universitatis Mosquеnsis (MW) / P. Smirnow. Pl. Armeniae ross. a. 1929. / Anoplanthus coccineus Walp. / ... / Daparuras / ... / 19.VI.1929 / N.ş 743 ..." (https://plant.depo.msu.ru/open/public/item/MW0723296 - MW 0723296.jpg - https://www.gbif.org/occurrence/1697840731; rev. Ó. Sánchez Pedraja Apr 2018); "Velmi výraznými prvky jsou zástupci zárazovitých (Orobanchaceae), např. nepřehlédnutelná rostlina s jediným červeným květem Phelypaea coccinea (obr. 10), která parazituje na chrpách (Centaurea)." (Ekrtová & Ekrt 2012: 70, 72, photo 10; accessed et rev. Ó. Sánchez Pedraja Jan 2015 [the photo seems to correspond to P. tournefortii]).

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Azerbaijan / Azərbaycan / Азәрбајҹан / Azerbaiyán / Aserbaidschan / Azerbaďdjan (Fischer 1852: 107, [Armenié, Bushe], sub Anoplanthus Tournefortii; Boissier & Buhse 1860: 169, sub Anoplanthus Tournefortii; Boissier 1879: 494, sub Anoplanthus coccineus var. peduncularis; Stapf 1915: 288, sub Phelypaea orientalis, flore coccinea; Beck 1930: 44, sub P. tournefortii; Karjagin 1957)

Khizi / Xızı Rayon / Хызинский район: 1. "Anoplon Bieber / steinii / ... 27 Juin 56 [1856?] // Armenia Rossica [Dübrar Dağı?, near Altıağac / Alty-Agatsch?] / Seidlitz [N.K. von Seidlitz] / Herb. Al. de Bunge". – 2. ibidem (https://science.mnhn.fr/institution/mnhn/collection/p/item/p02970939 - P 02970939.jpg - hb. E. Cosson - https://www.gbif.org/occurrence/439109406; rev. Ó. Sánchez Pedraja Nov 2011, sub D. tournefortii [three (2+1) specimens on the lower part of the sheet, close to Bunge's labels]); "Russian Armenia" (Stapf 1915: 288, sub Phelypaea orientalis, flore coccinea); "Calculated" (Domina & Raab-Straube 2010, sub P. tournefortii).

Khojavend / Xocavənd Rayon / Ходжавендский район [unconfirmed]: "Tale Dschagritschai, bei Takjalta Kulu [Dağdöşü / Tak] (Bayern!)" (Beck 1930: 44, sub P. tournefortii).

Nakhchivan Autonomous Republic / Naxçıvan Muxtar Respublikası / Нахичеванская Автономная Республика (Boissier & Buhse 1860: 169, sub Anoplanthus Tournefortii)

Бабекский район / Babek Rayon: "444. Anoplanthus Tournefortii / Walp. ex parte / Puskoh bei Nachitschewan / 31/V, 47 [31 May 1847] / Buhse [leg.] // Iter Persicum. / Buhse, 1847-1849 / Herb. Al. de Bunge" (https://science.mnhn.fr/institution/mnhn/collection/p/item/p02970936 - P 02970936.jpg - E. Cosson - https://www.gbif.org/occurrence/438443080; rev. Ó. Sánchez Pedraja Nov 2011, sub D. tournefortii [the two specimens on the upper left corner close to Buhse's label]); "Phelypaea coccinea (M. Bieb.) Poir. / Nakhichivan / 1847 / F. Buhse" (LD:General:2073901 - https://www.gbif.org/es/occurrence/2012691834 [n.v.]); "Bei Nachitschewan im Dschagrithal [Dzhagry / Cəhriçay / Джагричай, Babek District / Бабек] beim Dorfe Pusko, auf Pyrethrum achillaefolium (nicht auf P. myriophillum, wie Fischer Bull de la Soc. Imp. de Moscou 1852 nş I 107, angiebt), 31 Mai 1847 (florens). Nş 444" (Boissier & Buhse 1860: 169, sub Anoplanthus Tournefortii); "Armeniâ Rossicá (Buhse!)" (Boissier 1879: 494, sub Anoplanthus coccineus var. peduncularis); "Transcaucasia: Near Pusko in the Dshagri valley [Dzhagry / Cəhriçay], north of Nahitshewan [Nachitschewan / Nakhchivan / Нахичевань], on Pyrethrum myriophyllum, C.A. Mey., Buhse" (Stapf 1915: 293, sub P. tournefortii); "in der Provinz Nachitschevan [Nakhchivan Autonomous Republic] um Nachitschevan [Nakhchivan / Naxçıvan] bei Puskoh (Buhse!)" (Beck 1930: 44, sub P. tournefortii); "Herbarium Universitatis Mosquеnsis (MW) / ... / Filiatonis Azerbajdzhanicae Academiae Scientiarum URSS / Herbarium Azerbajdzhanicum / Phelураea coccinea Poir. / Transcaucasia, Azerbajdzhan, respublica Nachitshevan, distr. / Nachitshevan, prope pag. Buzgov [Aşağı Buzqov]. / in calcareis 29.V.1939 Leg. E. Chalivov / Det. ..." (https://plant.depo.msu.ru/open/public/item/MW0723302 - MW 0723302.jpg - https://www.gbif.org/occurrence/1697840776; rev. Ó. Sánchez Pedraja Apr 2018).

Садаракский район / Sadarak Rayon: "Herbarium Universitatis Mosquеnsis (MW) / Phelураea tournеfortii Desf. / Армения. / Вединский район [Vedi rayonu / Ararat rayonu] / В 2 км к югу от с. Кярки [2 km to the south of the village Karki / Kərki / Kyarki / Tigranashen / Тигранашен] можжевеловое редколесье, 1400 м. / 29 мая 1973 г. [29.05.1973] Н. Шведчикова [N. Shvedchikova]" (https://plant.depo.msu.ru/open/public/item/MW0723306 - MW 0723306.jpg - https://www.gbif.org/es/occurrence/1697840797; rev. Ó. Sánchez Pedraja Apr 2018, on Tanacetum sp.).

Шарурский район / Sharur Rayon: "Herbarium Universitatis Mosquеnsis / Azərbaycan Mərkəzi Herвarium /... / Phelураea Tournеfortii Desf. / Transcaucasia, respublica Naсhitschevan, prope p. / Karakush, in herbosis. / Leg L. Prilipkо / 31-V-1934 Det. L. Prilipkо" (https://plant.depo.msu.ru/open/public/item/MW0723308 - MW 0723308.jpg - https://www.gbif.org/occurrence/1697840815; rev. Ó. Sánchez Pedraja Apr 2018).

Unknown location: "Herbarium Universitatis Mosquеnsis (MW) / ... / Phelураea coccinea Poir. / ... Зривано-Нахичеванская ... [Кавказ, Азербайджан / Caucasus, Azerbaijan] / 10.VI.1931 / Leg. И.И. Карягин [I.I. Karyagin]" (https://plant.depo.msu.ru/open/public/item/MW0723298 - MW 0723298.jpg - https://www.gbif.org/occurrence/1697840765; rev. Ó. Sánchez Pedraja Apr 2018, sub D. tournеfortii).

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Georgia / Sakartvelo / Georgien / Géorgie / Грузия [needs be confirmed]

"Anoplon Biebersteinii C.A.M. / Caucas / 1850. / leg. Kovalevsky" (https://science.mnhn.fr/institution/mnhn/collection/p/item/p02970929 - P 02970929.jpg - hb. Drake - https://www.gbif.org/occurrence/439442856, rev. Ó. Sánchez Pedraja Nov 2011 [sub P. tournеfortii, only one specimen well preserved, "calyx cup-shaped with a broad upper lip which is shallowly threetoothed at the apex and a lower lip of two longer, more deeply cut lobes; scale-leaves concentrated towards the bases of the stems" (Cullen 2010: 384)]); "Georgia" (Tarkhnishvili & Chaladze 2013 in Georgian Biodiversity Database, accessed Dec 2010 and Oct 2017).

Note: The gathering of Hohenaker in May 1831 and 1834 seem to correspond to D. coccinea: "pratis montosis umbrosis Georgiae caucasicae (Hohenacker !)" (Reuter 1847: 42, sub Anoplanthus biebersteinii Reut.); "in pratis montosis Georgiae Caucasicae (Hoh!)" (Boissier 1879: 494, sub Anoplanthus coccineus var. peduncularis); "Georgien (Hohenacker nach Boissier)" (Beck 1930: 44, sub P. tournefortii).

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Iran / ایران‎ / Irán [Persia] [needs be confirmed (see P. boissieri)] (Gardner & Gardner 2014: 119, sub D. tournefortii)

West Azerbaijan Province: (https://gramhir.com/media/2587342286867981834 by @hadi.urmulu, sub P. coccinea; accessed et rev. Ó. Sánchez Pedraja Sep 2021 [sub P. tournefortii, apparently on Tanacetum sp.]).

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Turkey / Türkiye / Tuquía / Turkije (Reuter 1847: 43, sub Anoplanthus tournefortii; Boissier 1879: 494, sub Anoplanthus coccineus var. peduncularis; Stapf 1915: 293, sub P. tournefortii; Beck 1930: 44, sub P. tournefortii; Bakis & al. Distribution of The Taxon Over Turkey in Turkish Plants Data Service (TÜBİVES) (accessed Sep 2017); Bizim Bitkiler 2021 (http://www.bizimbitkiler.org.tr...id=9705, sub P. tournefortii); Piwowarczyk & al. 2019: 18, sub D. tournefortii, epitype of P. tournefortii)

Eastern Anatolia Region (Doğu Anadolu Bölgesi) "Phelypaea tournefortii: IMG_3551, IMG_3555, IMG_3579 [on Tanacetum sp.] (photos by Dana Cromie in Eastern Turkey, May 2011 [https://www.flickr.com]; accessed Feb 2013)

Northeast Anatolia Region / Kuzeydoğu Bölgesi

Ağrı Province: "B9 Ağri: Murat valley, 1600 m, D. 44038!" (Gilli 1982: 2, sub P. tournefortii).

Bayburt Province / Bayburt ili: "Huet de Pavillon / Plantae Orientales exsiccatae. / Anoplanthus Tournefortii  Walpers. / Reut. In D.C. Prod. / Im valle Kapuclu [Kussuklu / Ch'orokhi / Çoruh / Tchorokh?] 4-5000 P S. m / Inter Baiburt [Bayburt] et Erzeroum [Erzurum] / Maio 1853  Armenia" (https://science.mnhn.fr/institution/mnhn/collection/p/item/p02970936 - P 02970936.jpg - E. Cosson - https://www.gbif.org/occurrence/438443080; rev. Ó. Sánchez Pedraja Nov 2011, sub D. tournefortii [only one specimen on the lover left corner close to Huet de Pavillon's label]); 1. "Huet de Pavillon / Plantae Orientales exsiccatae. / Anoplanthus Tournefortii  Walpers. / Reut. In D.C. Prod. / Im valle Kapuclu [Kussuklu / Ch'orokhi / Çoruh / Tchorokh?] 4-5000 P S. m / Inter Baiburt [Bayburt] et Erzeroum [Erzurum] / Maio 1853  Armenia". – 2. "Herb. Mus. Paris / Anoplanthus / Phelipaea Tournefortii / Desf. / Armenie. / Collection de Mr. Simon. / donnée par Mr. le Cte. de Challaye 1855" (https://science.mnhn.fr/institution/mnhn/collection/p/item/p02970941 - P 02970941.jpg - https://www.gbif.org/occurrence/438443082; rev. Ó. Sánchez Pedraja Nov 2011, epitype of P. tournefortii (Piwowarczyk & al. 2019: 18, sub D. tournefortii)); "Bayburt" (Distribution of The Taxon Over Turkey in Turkish Plants Data Service (TÜBİVES) Version 2.0 BETA; accessed Sep 2017).

Erzurum Province [Western Armenia]: Armenia (Desfontaines 1807: t. 21; 1808: 16, t. 10 and https://science.mnhn.fr/institution/mnhn/collection/p/item/p02970942 - P 02970942.jpg - https://www.gbif.org/occurrence/437111828, sub Phelypaea tournefortii); "native of Armenia" (Stapf 1915: 285, 288, sub Phelypaea orientalis, flore coccinea); "H. Native of Armenia" (Don, 1838: 633, sub Anoplon tournefortii); "In Armeniâ (Tournef.)" (Reuter 1847: 43, sub Anoplanthus tournefortii); "Huet de Pavillon / Plantae Orientales exsiccatae. / Anoplanthus Tournefortii  Walpers. / Reut. In D.C. Prod. / Im valle Kapuclu [Kussuklu / Ch'orokhi / Çoruh / Tchorokh?] 4-5000 P S. m / Inter Baiburt [Bayburt] et Erzeroum [Erzurum] / Maio 1853  Armenia" (https://science.mnhn.fr/institution/mnhn/collection/p/item/p02970936 - P 02970936.jpg - E. Cosson - https://www.gbif.org/occurrence/438443080, rev. Ó. Sánchez Pedraja Nov 2011 [only one specimen on the lover left corner close to Huet de Pavillon's label]); 1. idem. – 2. "Herb. Mus. Paris / Anoplanthus / Phelipaea Tournefortii / Desf. / Armenie. / Collection de Mr. Simon. / donnée par Mr. le Cte. de Challaye 1855" (https://science.mnhn.fr/institution/mnhn/collection/p/item/p02970941 - P 02970941.jpg - https://www.gbif.org/occurrence/438443082, rev. Ó. Sánchez Pedraja Nov 2011); 1. "Anoplon Tournefortii / ou / Phelipaea coccinea". – 2. " 55. Top Dagh pres [?] / Erzeroum [Erzurum], Arménie. / Simon, 1854" (https://science.mnhn.fr/institution/mnhn/collection/p/item/p02970940 - P 02970940.jpg - E. Cosson - https://www.gbif.org/occurrence/437444715, rev. Ó. Sánchez Pedraja Nov 2011 [seven specimes well preserved]); 1. "Orobanchea / Anoplon Tournefortii / Syn.:? / Phelipaea coccinea?". – 2. "Herb. Mus. Paris / Anoplanthus / Phelipaea Tournefortii / Desf. / Armenie. / Collection de Mr. Simon. / donnée par Mr. le Cte. de Challaye 1855" (https://science.mnhn.fr/institution/mnhn/collection/p/item/p02970943 - P 02970943.jpg - https://www.gbif.org/occurrence/438443081; rev. Ó. Sánchez Pedraja Nov 2011); "in Armeniâ Turcicâ circa Erzerum [Erzurum / Karin, Anatolia, terra class. of P. tournefortii Desf.] (Tourn!, Huet!, Calv!)" (Boissier 1879: 494, sub Anoplanthus coccineus var. peduncularis); "In Kurdistaniâ [Turkish Kurdistan] (J. Brant! Strangways in h. Benth.!)" (Reuter 1847: 43, sub Anoplanthus tournefortii); "Kurdistaniâ (Barré de Lancy [vice-consul de France ŕ Mersine et Adana])!" (Boissier 1879: 494, sub Anoplanthus coccineus var. peduncularis); "Herbier de Ch. d'Alleizette / Plantes ... levant d'échanges [?] / Phelypaea coccinea Pers. / (Anoplanthus Tournefortii) / Asie Mre [Mineure] env. Erzurum / Juillet 1904" (https://data.biodiversitydata.nl/naturalis/specimen/L.2821536 - L 2821536.jpg - https://www.gbif.org/es/occurrence/2516614582; rev. Ó. Sánchez Pedraja Oct 2017); "discovered in Armenia, probably in the neighbourhood of Erzurum by Tournefort in 1701 [June 15 to July 6]" (Stapf 1915: 285, 289, 293, sub Phelypaea orientalis, flore coccinea and sub P. tournefortii); "Turkish Kurdistan" (Stapf 1915: 288, sub Phelypaea orientalis, flore coccinea); "Turkish Armenia: Upper Chorok valley, near Masut [Çakılı], E. of Baibut [Baiburt / Bayburt], 1200-1800 m, Huet de Pavillon! Near Erzurum, Zohrab, 826! Calver and Zohrab, 158! Kurdistan; without precise locality, Brant! Barre de Lancy (Hb. Gaillardot, 2555)" (Stapf 1915: 293, sub P. tournefortii); "ad radices Pyrethri myriophylli C.A. Meyer [Tanacetum polycephalum subsp. duderanum (Boiss.) Podlech] et Achilleae spec. ... Armenien: Provinz Erzerum (Tournefort, Calvert nach Boissier); im Tale Kussuklu zwischen Baiburt und Erzerum bei 4000-5000' (Huet de Pavillon!) ... Kurdistan (Barre de Lancy nach Boissier)" (Beck 1930: 44, sub P. tournefortii); "E. Anatolia. A8 Erzurum: Sari Koma pass, 2600 m, Furse & Synge 850! Gümüşane: Kussuklu valley, Bayburt to Erzurum, 1220-1525 m, Huet." (Gilli 1982: 2, sub P. tournefortii); "https://www.ncbi.nlm.nih.gov/nuccore/AY209285: Turkey, NE Anatolia, between Bayburt and Erzurum, between the village Maden and Kop Geçidi, 1840 m, N 40° 04' 49'' E 40° 25' 34'', on Tanacetum sp. (Asteraceae), G. Schneeweiss & C.-G. Jang (WU: S&T 7764 [http://www.parasiticplants.siu.edu/Orobanchaceae/images/Diphelypaea_tourn_habit.jpg]) (Carlón & al. 2005: 63 sub P. tournefortii Desf.)"; "Gümüşane, Mescit Daglari: an der Straße zwischen Bayburt und Erzurum, zwischen dem Ort Maden und dem (Pass) Kop Geçidi [Bayburt Province]; 40°04'49'' N, 40°25'34'' E. 1840 m, Schneeweiß, G. & Jang, C.-G. 7764 - https://www.ncbi.nlm.nih.gov/nuccore/AY209285, 2002-06-15" (WU 0032431 - https://wu.jacq.org/WU0032431 - https://www.jacq.org/detail.php?ID=39026); "Phelypaea tournefortii Desf. / Erzurum: Karaçoban, Kırmızı Tuzla surroundings, steppe, 03 vi 2006, 39° 17' 377'' N 42° 06' 329'' E, 1592 m, LFM 770, Ir.-Tur., Hc. On Centaurea schischkinii [erroneous host-plant?]" (Behçet & al. 2009: 147, sub P. tournefortii); "Erzurum" (photo 69109 by Hayati Keser in www.treknature.com, accessed Mar 2012); "Cat. 2, 1e prijs, Rinus Bode, Phelypaea tournefortii, Kaskargebergte [Pontisch Gebergte / Kaçkar Dağları / Pontic Mountains / Kuzey Anadolu Dağları / North Anatolian Mountains], N-O Turkije" (photo 01 by NRV - Nederlandse Rotsplanten Vereniging - fotowedstrijd van 2012; accessed Feb 2013); "Fotografováno dne 8.6.2013 (Turecko, oblast Aşkale [Erzurum Province], na úbočí Kop Geçidi [Bayburt Province])" (https://botany.cz/cs/phelypaea-tournefortii/ by Pavla Dvorská-Hochmanová, sub P. tournefortii, Autor Eva Rencová, 30.6.2013 in Botany.cz; 2007-2014; accessed Jan 2015); "Narman" (https://gramhir.com/media/2379109340939464334, https://gramhir.com/media/2327631659157093838 and https://gramhir.com/media/2528694368643309541 by @doganinsirdaslari, sub P. coccinea; accessed et rev. Ó. Sánchez Pedraja Aug 2021 [sub P. tournefortii, apparently on Tanacetum sp.]).

Kars Province: "im Distr. Kaghyzman [Kagizman / Kağızman (Armenian Highlands)] am Sattel Kara dagh (S. Michailowsky)" (Beck 1930: 44, sub P. tournefortii); "A9 Kars: Yağmurlu Da. [Mount Yagmurlu], 2200 m, D. 30822!" (Gilli 1982: 2, sub P. tournefortii); Phelipaea tournefortii Desf., Prov. Kars: Yaqmurlu Daq between Karaurgan & Sa, 2200 m, 13.06.1957, Davis, P.H., Hedge, J.C. 29480 (http://specimens.kew.org/herbarium/27974.000 - https://www.gbif.org/occurrence/912025203 [n.v.], https://data.rbge.org.uk/herb/E00227405 - https://www.gbif.org/occurrence/1228038038 [n.v., sub Anoplanthus coccineus] et https://data.rbge.org.uk/herb/E00222695 - https://www.gbif.org/occurrence/1228044359 [n.v., sub Anoplanthus coccineus]).

Central East Anatolia Region / Ortadoğu Anadolu Bölgesi

Malatya Province / Malatya ili: "Malatya dolaylarında çekilmiştir [it was taken near Malatya]" (photos http://yabanicicekler.com/flower/phelypaea-coccinea-53 [1/3-2/3-3/3 (53)] sub Phelypaea coccinea in Türkiye'nin Çiçekleri - Yabani Çiçekler; accessed et rev. Ó. Sánchez Pedraja Oct 2017 [the three photos seem to correspond to P. tournefortii ("Stems and leaves covered with dense to sparse, long, multicellular white hairs; calyx cup-shaped with a broad upper lip which is shallowly threetoothed at the apex and a lower lip of two longer, more deeply cut lobes" (Cullen 2010: 384)), moreover we can observe the host-plant [Tanacetum sp.] in the photo 2/3]).

Tunceli Province / Tunceli ili [formerly Dersim Province]: "Flora Turcomanniae Asiaticae / Iter Leydense 1959 / Collectores: E. Hennipman, P. Nijhoff, C. Swennen, A.S.Tulp, / W.J.M. Vader, V.J.J.O. de Wildee / No. 1635 Date: 29-5 Altitude ± : 2000 m. / Prov.: Tunçeli. Locality: ± 5 km N. of Pülümür (along the road Elâzig-Erzincan) / ... / Flowers humid neglected field. Clayish soil. / Flowers bright red. / (Host Centaurea ... / Anoplanthus coccinus Walp." (https://data.biodiversitydata.nl/naturalis/specimen/L.2825068 - L 2825068.jpg - https://www.gbif.org/occurrence/2514627751; rev. Ó. Sánchez Pedraja Jun 2021 [sub P. tournefortii, plants with scales only at the base of the hairy stem and calyx with conspicuous hairs; the indicated host-plant (Centaurea) is apparently erroneous but see P. boissieri]); idem (https://data.biodiversitydata.nl/naturalis/specimen/WAG.1141086 - WAG 1141086.jpg - https://www.gbif.org/occurrence/2515854856; rev. Ó. Sánchez Pedraja Jun 2021 [sub P. tournefortii but see P. boissieri]); "Tunceli, S¸Y 3245" (Zare & al. 2014, sub D. tournefortii); "Karagöz, Pülümür [39°26'49.8"N 39°40'05.0"E]", 2 jul 2015 (https://www.inaturalist.org/observations/38100719 - https://www.gbif.org/es/occurrence/2563645191 by Mustafa Gökmen, sub Diphelypaea coccinea, in iNaturalist; accessed et rev. Ó. Sánchez Pedraja Feb 2020 [sub P. tournefortii, apparently on Tanacetum sp.]).

Van Province: "Coteaux secs, entre Van et Hokotz-vank ['Hokvotz-vank, monastére isolé, á environ 45 kilom. au sud-sudest de Van, tout prčs et au sud-ouest de Soussouz-dagh'.] (exs. 110)" (Aznavour 1917: 28, sub Anoplanthus coccineus var. peduncularis Boiss.); "zwischen Van und Hokotz-vank (D. Tchitouny nach Aznavour 1917)" (Beck 1930: 44, sub P. tournefortii); "Van: between Van and Hokotz church, Tchitouny" (Gilli 1982: 2, sub P. tournefortii); "Bild 117 - Phelypaea toumefortii (Orobanchaceae) auf Tanacetum schmarotzend, am Fuß eines felsigen Hanges nördlich von Muradiye; etwa natürliche Größe." (Sorger 1994: 174, sub P. tournefortii); "June 13, 2002. Near Caldiran [Çaldıran, Van], eastern Turkey" (sub P. coccinea, 2006 Old Dominion University, accessed Dec 2010); Van: steppes le long du chemin d’accčs aux chutes de Muradiye, 1826 m, UTM 38S 0392562 / 4324169, leg. P. Coulot & al., 24.05.2009 [habit, flower, calyx, scale-leaves]; Lake Van / Phelypaea tournefortii - parasite on Tanacetum species (http://www.fands.org.uk/FANDS_images/VAN/_0012090.jpg in http://www.fands.org.uk; accessed Feb 2013); "Sommerwurz in der Osttürkei nahe der iranischen Grenze", June 1990 (https://commons.wikimedia.org/wiki/File:Phelypaea_tournefortii_1.jpg by Ziegler175; accessed Feb 2020).

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References.

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Bakis, Y., Babac, M.T., & Uslu, E. 2011. Updates and improvements of Turkish Plants Data Service (TÜBİVES). In Health Informatics and Bioinformatics (HIBIT), 2011 6th International Symposium on (pp. 136-140). IEEE. TÜBİVES. Türkiye Bitkileri Veri Servisi [Turkish Plants Data Service]. URL: http://194.27.225.161/yasin/tubives/index.php - http://194.27.225.161/yasin/tubives/index.php?sayfa=1&tax_id=7340 (accessed Sep 2017).

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Behçet, B., Özgökçe, F., Ünal, M. & ve Karabacak, Ö. 2009. The flora of Kırmızı Tuzla (Karaçoban, Erzurum/Turkey) and Bahçe Tuzlası (Malazgirt, Mus/Turkey) and their environment. Biological Diversity and Conservation 2(3): 122-155. URL: https://www.researchgate.net/publication/274721110 - https://www.academia.edu/30756558.

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